Potentilla millefolia |
Potentilla subvahliana |
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cut-leaf cinquefoil, feather cinquefoil, feather or many-leaf or Klamath cinquefoil, many leaf cinquefoil |
high arctic cinquefoil |
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Habit | Plants ± rosetted; taproots fleshy-thickened. | Plants usually cushion-forming. |
Caudex branches | stout, columnar, sheathed with marcescent whole leaves. |
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Stems | usually prostrate, sometimes ± decumbent, 0.4–2(–3) dm, lengths 1–2 times basal leaves. |
erect, 0.2–0.6(–1.1) dm, lengths 2–4 times basal leaves. |
Basal leaves | pinnate with distal leaflets ± confluent, 2–15(–20) × 1–3 cm; petiole 0.5–2(–3) cm, straight hairs sparse to abundant, appressed to spreading, 0.5–1.5 mm, stiff, cottony hairs absent, glands absent or sparse; primary lateral leaflets (3–)5–13 per side, on distal 2/3–3/4+ of leaf axis, separate to overlapping, largest ones cuneate to flabellate, 0.5–1.5(–2) × 0.5–2 cm, distal 2/3 to whole margin palmately or unevenly, rarely pinnately, incised 2/3 to completely to midvein, ultimate teeth or segments (1–)2–10, linear to broadly oblanceolate, 2–10 × (0.5–)1–2 mm, apical tufts to 1 mm, surfaces green to grayish green, not glaucous, straight hairs sparse to abundant, appressed to spreading, 0.5–1.5(–2) mm, stiff, cottony hairs absent, glands sparse to common. |
0.5–2.5(–3) cm; petiole 0.3–1.5(–2) cm, long hairs common to abundant, spreading to ascending, 1(–2) mm, ± weak, rarely stiff, smooth, crisped/short-cottony hairs usually absent, sometimes sparse, glands sparse to common; leaflets separate to slightly overlapping, central obovate, 0.5–1.2(–1.5) × 0.4–0.9(–1) cm, subsessile to short-petiolulate, base cuneate, margins revolute, distal 1/2 incised 1/2–3/4 to midvein, teeth (1–)2–3(–4) per side, ± distant, surfaces ± dissimilar, abaxial grayish or yellowish white, long hairs 0.5–1 mm, cottony-crisped hairs ± dense (often obscured by long hairs), adaxial dark green to greenish gray, long hairs ± abundant, other hairs absent. |
Cauline leaves | (0–)1–2. |
(0–)1(–2). |
Inflorescences | 3–6(–10)-flowered, loosely cymose, sometimes racemiform. |
1(–2)-flowered. |
Pedicels | (0.5–)1–2(–4.5) cm, ± recurved in fruit. |
1–2(–3) cm in flower, to 5.5 cm in fruit. |
Flowers | epicalyx bractlets ± elliptic, 2–4(–6) × 1–2(–2.5) mm; hypanthium 3–6 mm diam.; sepals 4–6(–8) mm, apex acute; petals 4–8(–10) × 3–7(–9) mm; filaments 2–3.5 mm, anthers 0.7–1 mm; carpels 10–30, styles (1.5–)2–3 mm. |
epicalyx bractlets lanceolate to elliptic-ovate, rarely linear, 3–6(–7) × 0.8–2(–2.3) mm, (1/2–)2/3 to nearly as wide as sepals, margins flat or ± revolute, red glands absent; hypanthium 2.5–4 mm diam.; sepals 3–5(–6) mm, apex obtuse to subacute; petals 4–8(–9) × 4–9 mm, longer than sepals; filaments 1–1.3 mm, anthers 0.3–0.5 mm; carpels 25–35, apical hairs absent, styles conic-columnar, ± papillate-swollen on less than proximal 1/5, 0.9–1.1 mm. |
Achenes | 1.5–2 mm, smooth, often ± carunculate. |
1.2–1.6 mm. |
2n | = 28 (Russian Far East). |
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Potentilla millefolia |
Potentilla subvahliana |
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Phenology | Flowering spring–summer. | Flowering late spring to summer. |
Habitat | Vernally to permanently wet meadows, moist openings in conifer forests and sagebrush, alkaline flats | Rocky alpine heaths, rock outcrops and crevices, scree and talus, dry tundra, coastal bluffs, stabilized sand dunes, usually on calcareous bedrock |
Elevation | 700–2200 m (2300–7200 ft) | 0–1700 m (0–5600 ft) |
Distribution |
CA; NV; OR
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AK; AB; BC; NT; NU; QC; YT; Greenland; e Asia (Russian Far East) |
Discussion | Potentilla millefolia occurs from central Oregon to the east side of the Sierra Nevada of California, with a disjunct occurrence on the alkaline flats of Reese River Valley, Nevada. Significant variation occurs in vestiture type, leaflet dissection, and flower size, but with minimal geographic correlation. The most distinctive variant, represented by the type of P. klamathensis, has relatively long, slender, spreading, pustule-based hairs, often intermixed with shorter hairs. This vestiture type does not appear to be correlated with any other characters or geographic distribution and may vary within a population. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
The major part of the range assigned to Potentilla vahliana by E. Hultén (1968) belongs to P. subvahliana. Two morphologic types are present within what is accepted here as Potentilla subvahliana. Plants corresponding to the type of P. subvahliana (Wrangel Island) are widespread throughout northeastern Asia (Chukotka) and arctic North America to northwestern Greenland. Some plants in alpine Alaska, Yukon, and east to Amundsen Gulf, Nunavut, are redder, more densely columnar, and less hairy, with smaller leaves having fewer and narrower lobes, more slender one-flowered stems, narrow and entire bracts, narrower sepals, and much narrower epicalyx bractlets. Potentilla subvahliana is tetraploid, perhaps an allopolyploid, and could consist of lineages from different parental combinations. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 9, p. 174. | FNA vol. 9, p. 203. |
Parent taxa | Rosaceae > subfam. Rosoideae > tribe Potentilleae > Potentilla > sect. Multijugae | Rosaceae > subfam. Rosoideae > tribe Potentilleae > Potentilla > sect. Niveae |
Sibling taxa | ||
Synonyms | P. klamathensis, P. millefolia var. klamathensis, P. plattensis var. klamathensis, P. plattensis var. millefolia | |
Name authority | Rydberg: Bull. Torrey Bot. Club 23: 433, plate 277, figs. 1–5. (1896) | Jurtzev: in A. I. Tolmatchew, Fl. Arct. URSS 9(1): 319. (1984) |
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