Potentilla millefolia |
Potentilla stipularis |
|
---|---|---|
cut-leaf cinquefoil, feather cinquefoil, feather or many-leaf or Klamath cinquefoil, many leaf cinquefoil |
stipulate cinquefoil, stipulated cinquefoil |
|
Habit | Plants ± rosetted; taproots fleshy-thickened. | |
Glands | mostly absent. |
|
Stems | usually prostrate, sometimes ± decumbent, 0.4–2(–3) dm, lengths 1–2 times basal leaves. |
(0.2–)1–2.5(–3.5) dm. |
Basal leaves | pinnate with distal leaflets ± confluent, 2–15(–20) × 1–3 cm; petiole 0.5–2(–3) cm, straight hairs sparse to abundant, appressed to spreading, 0.5–1.5 mm, stiff, cottony hairs absent, glands absent or sparse; primary lateral leaflets (3–)5–13 per side, on distal 2/3–3/4+ of leaf axis, separate to overlapping, largest ones cuneate to flabellate, 0.5–1.5(–2) × 0.5–2 cm, distal 2/3 to whole margin palmately or unevenly, rarely pinnately, incised 2/3 to completely to midvein, ultimate teeth or segments (1–)2–10, linear to broadly oblanceolate, 2–10 × (0.5–)1–2 mm, apical tufts to 1 mm, surfaces green to grayish green, not glaucous, straight hairs sparse to abundant, appressed to spreading, 0.5–1.5(–2) mm, stiff, cottony hairs absent, glands sparse to common. |
3–7(–10) cm; petiole 2–5(–7.5) cm, hairs absent or sparse, appressed to spreading, 0.5–0.8 mm, stiff, glands absent or nearly so; leaflets 7–11, central one oblanceolate-elliptic, 1–2(–3) × 0.3–0.8 cm, distal 1/4 or less of margin incised less than 1/10(–1/4) to midvein, teeth 1–2(–5) per side, surfaces similar, abaxial pale green to reddish, often glaucous, hairs absent or sparse to common on primary veins, appressed to ascending, 0.6–1 mm, stiff, adaxial green or reddish, glabrous or nearly so. |
Cauline leaves | (0–)1–2. |
1–3; stipules fused with all or most of petiole, free portion shorter than fused portion. |
Inflorescences | 3–6(–10)-flowered, loosely cymose, sometimes racemiform. |
2–5-flowered. |
Pedicels | (0.5–)1–2(–4.5) cm, ± recurved in fruit. |
1–3 cm (proximalmost to 6 cm). |
Flowers | epicalyx bractlets ± elliptic, 2–4(–6) × 1–2(–2.5) mm; hypanthium 3–6 mm diam.; sepals 4–6(–8) mm, apex acute; petals 4–8(–10) × 3–7(–9) mm; filaments 2–3.5 mm, anthers 0.7–1 mm; carpels 10–30, styles (1.5–)2–3 mm. |
epicalyx bractlets lanceolate to narrowly lanceolate, 3–5 × 0.7–1(–1.5) mm; hypanthium 2.5–3 mm diam.; sepals 4–6 mm, apex acute; petals (4–)6–8 × 4–6 mm; filaments 1.6–2 mm, anthers 0.4–0.6 mm; carpels 40–50, styles 0.7–0.9 mm. |
Achenes | 1.5–2 mm, smooth, often ± carunculate. |
1.1–1.3 mm, smooth. |
2n | = 28 (Eurasia). |
|
Potentilla millefolia |
Potentilla stipularis |
|
Phenology | Flowering spring–summer. | Flowering summer. |
Habitat | Vernally to permanently wet meadows, moist openings in conifer forests and sagebrush, alkaline flats | Herb meadows, alluvial meadows and flats, open shrub thickets, Salix-Dryas tundra |
Elevation | 700–2200 m (2300–7200 ft) | 10–600 m (0–2000 ft) |
Distribution |
CA; NV; OR
|
AK; Greenland; Eurasia |
Discussion | Potentilla millefolia occurs from central Oregon to the east side of the Sierra Nevada of California, with a disjunct occurrence on the alkaline flats of Reese River Valley, Nevada. Significant variation occurs in vestiture type, leaflet dissection, and flower size, but with minimal geographic correlation. The most distinctive variant, represented by the type of P. klamathensis, has relatively long, slender, spreading, pustule-based hairs, often intermixed with shorter hairs. This vestiture type does not appear to be correlated with any other characters or geographic distribution and may vary within a population. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
The occurrences of Potentilla stipularis in Alaska are a continuation of the range from Asia. The six known occurrences in eastern and northeastern Greenland are disjunct (A. E. Porsild 1964) and were assigned to var. groenlandica by Sørensen, differing from Asian and Alaskan counterparts mainly in quantitative features: 9–11 leaflets, 7–11 teeth per leaflet, and not conspicuously glaucous (G. Halliday, pers. comm.). Here var. groenlandica is considered a high-arctic ecotype that provisionally is not accepted taxonomically. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 9, p. 174. | FNA vol. 9, p. 147. |
Parent taxa | Rosaceae > subfam. Rosoideae > tribe Potentilleae > Potentilla > sect. Multijugae | Rosaceae > subfam. Rosoideae > tribe Potentilleae > Potentilla > sect. Chrysanthae |
Sibling taxa | ||
Synonyms | P. klamathensis, P. millefolia var. klamathensis, P. plattensis var. klamathensis, P. plattensis var. millefolia | P. stipularis var. groenlandica |
Name authority | Rydberg: Bull. Torrey Bot. Club 23: 433, plate 277, figs. 1–5. (1896) | Linnaeus: Sp. Pl. 1: 498. (1753) |
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