Potentilla millefolia |
Potentilla rubricaulis |
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cut-leaf cinquefoil, feather cinquefoil, feather or many-leaf or Klamath cinquefoil, many leaf cinquefoil |
red-stem cinquefoil, Rocky Mountain cinquefoil |
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Habit | Plants ± rosetted; taproots fleshy-thickened. | |
Caudex branches | not sheathed with marcescent whole leaves. |
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Stems | usually prostrate, sometimes ± decumbent, 0.4–2(–3) dm, lengths 1–2 times basal leaves. |
ascending to nearly erect, 1.5–4 dm. |
Basal leaves | pinnate with distal leaflets ± confluent, 2–15(–20) × 1–3 cm; petiole 0.5–2(–3) cm, straight hairs sparse to abundant, appressed to spreading, 0.5–1.5 mm, stiff, cottony hairs absent, glands absent or sparse; primary lateral leaflets (3–)5–13 per side, on distal 2/3–3/4+ of leaf axis, separate to overlapping, largest ones cuneate to flabellate, 0.5–1.5(–2) × 0.5–2 cm, distal 2/3 to whole margin palmately or unevenly, rarely pinnately, incised 2/3 to completely to midvein, ultimate teeth or segments (1–)2–10, linear to broadly oblanceolate, 2–10 × (0.5–)1–2 mm, apical tufts to 1 mm, surfaces green to grayish green, not glaucous, straight hairs sparse to abundant, appressed to spreading, 0.5–1.5(–2) mm, stiff, cottony hairs absent, glands sparse to common. |
often both ternate and palmate on same plant, rarely subpalmate, 4–10 cm; petiole 2.5–7 cm, long hairs sparse to common, loosely appressed to ascending-spreading, 1–2 mm, ± weak to stiff, verrucose, short and/or ± crisped hairs common to abundant, cottony hairs absent, glands usually sparse; leaflets 3–5, proximalmost separated by 0(–1) mm, central oblong to obovate, 1.5–4 × 1–2.5 cm, petiolules 0–5 mm, distal 2/3–3/4 of margin incised 1/2–3/4, rarely +, to midvein, teeth (4–)5–8 per side, 4–5 mm, apical tufts 1 mm, abaxial surfaces gray to grayish white, long hairs abundant, cottony-crisped hairs usually dense, short hairs and glands absent or obscured, adaxial green to grayish green, long hairs sparse to common, 0.5–1.5 mm, stiff, short hairs absent or sparse, rarely common, crisped and cottony hairs absent, glands absent or sparse, rarely common. |
Cauline leaves | (0–)1–2. |
2–3. |
Inflorescences | 3–6(–10)-flowered, loosely cymose, sometimes racemiform. |
4–20-flowered, open, branch angle (10–)20–45°. |
Pedicels | (0.5–)1–2(–4.5) cm, ± recurved in fruit. |
0.5–3 cm, proximal to 5 cm. |
Flowers | epicalyx bractlets ± elliptic, 2–4(–6) × 1–2(–2.5) mm; hypanthium 3–6 mm diam.; sepals 4–6(–8) mm, apex acute; petals 4–8(–10) × 3–7(–9) mm; filaments 2–3.5 mm, anthers 0.7–1 mm; carpels 10–30, styles (1.5–)2–3 mm. |
epicalyx bractlets linear to narrowly lanceolate, 3–4 × 0.8–1.2 mm; hypanthium 4–6 mm diam.; sepals 4–5 mm, apex subacute to acute, glands usually ± sparse, not obscured; petals pale yellow, not overlapping, 5–7 × (4–)5–6.5 mm, distinctly longer than sepals; filaments 0.5–1.5 mm, anthers 0.4 mm; carpels 30–60, styles 0.9–1.1 mm. |
Achenes | 1.5–2 mm, smooth, often ± carunculate. |
1.2 mm. |
2n | = 56. |
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Potentilla millefolia |
Potentilla rubricaulis |
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Phenology | Flowering spring–summer. | Flowering summer. |
Habitat | Vernally to permanently wet meadows, moist openings in conifer forests and sagebrush, alkaline flats | Sandy lake and stream shores, open sandy forests, dry grassy slopes, sandy and loamy bluffs, rock crevices, scree |
Elevation | 700–2200 m (2300–7200 ft) | 0–1600 m (0–5200 ft) |
Distribution |
CA; NV; OR
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AK; AB; BC; NT; SK; YT
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Discussion | Potentilla millefolia occurs from central Oregon to the east side of the Sierra Nevada of California, with a disjunct occurrence on the alkaline flats of Reese River Valley, Nevada. Significant variation occurs in vestiture type, leaflet dissection, and flower size, but with minimal geographic correlation. The most distinctive variant, represented by the type of P. klamathensis, has relatively long, slender, spreading, pustule-based hairs, often intermixed with shorter hairs. This vestiture type does not appear to be correlated with any other characters or geographic distribution and may vary within a population. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
As addressed by B. Ertter et al. (2013), the name Potentilla rubricaulis is here restricted to relatively large plants with open inflorescences occurring mainly in glaciated parts of subarctic northwestern Canada and Alaska. Plants from the Chugach Mountains of southern Alaska tend to be more conspicuously glandular than elsewhere. The distinction between Potentilla rubricaulis and large forms of P. arenosa with supernumerary leaflets is problematic. Although both species have somewhat similar petiole vestiture (long, straight, verrucose hairs and a layer of short, stiff, or curly hairs), the latter species tends to have more stiffly spreading petiole hairs and prominently petiolulate central leaflets. The octoploid chromosome count (P. M. Dansereau and E. Steiner 1956) from Great Bear Lake area, Northwest Territories, probably belongs to Potentilla rubricaulis in the narrow sense, since that is its type locality. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 9, p. 174. | FNA vol. 9, p. 207. |
Parent taxa | Rosaceae > subfam. Rosoideae > tribe Potentilleae > Potentilla > sect. Multijugae | Rosaceae > subfam. Rosoideae > tribe Potentilleae > Potentilla > sect. Rubricaules |
Sibling taxa | ||
Synonyms | P. klamathensis, P. millefolia var. klamathensis, P. plattensis var. klamathensis, P. plattensis var. millefolia | P. dissecta var. rubricaulis, P. nivea subsp. rubricaulis |
Name authority | Rydberg: Bull. Torrey Bot. Club 23: 433, plate 277, figs. 1–5. (1896) | Lehmann: Nov. Stirp. Pug. 2: 11. (1830) |
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