Potentilla millefolia |
Potentilla modesta |
|
---|---|---|
cut-leaf cinquefoil, feather cinquefoil, feather or many-leaf or Klamath cinquefoil, many leaf cinquefoil |
modest cinquefoil |
|
Habit | Plants ± rosetted; taproots fleshy-thickened. | |
Caudex branches | usually not sheathed with marcescent whole leaves. |
|
Stems | usually prostrate, sometimes ± decumbent, 0.4–2(–3) dm, lengths 1–2 times basal leaves. |
ascending to nearly erect, (0.3–)0.5–1.5(–2.5) dm. |
Basal leaves | pinnate with distal leaflets ± confluent, 2–15(–20) × 1–3 cm; petiole 0.5–2(–3) cm, straight hairs sparse to abundant, appressed to spreading, 0.5–1.5 mm, stiff, cottony hairs absent, glands absent or sparse; primary lateral leaflets (3–)5–13 per side, on distal 2/3–3/4+ of leaf axis, separate to overlapping, largest ones cuneate to flabellate, 0.5–1.5(–2) × 0.5–2 cm, distal 2/3 to whole margin palmately or unevenly, rarely pinnately, incised 2/3 to completely to midvein, ultimate teeth or segments (1–)2–10, linear to broadly oblanceolate, 2–10 × (0.5–)1–2 mm, apical tufts to 1 mm, surfaces green to grayish green, not glaucous, straight hairs sparse to abundant, appressed to spreading, 0.5–1.5(–2) mm, stiff, cottony hairs absent, glands sparse to common. |
usually palmate, rarely also ternate on same plant or subpalmate, 2–6(–8) cm; petiole 0.5–3.5(–5) cm, long hairs common to abundant, ± ascending to almost spreading, 1–2 mm, weak to ± stiff, scarcely to ± verrucose, short and/or crisped hairs common to abundant, cottony hairs absent, glands sparse to common; leaflets (3–)5, proximalmost separated by 0–1 mm, central obovate to oblanceolate-elliptic, 0.5–2(–2.5) × 0.5–1(–1.5) cm, petiolules 1 mm, distal 3/4 to nearly whole margin incised 1/2–3/4 to midvein, teeth 2–5 per side, (1–)2–5 mm, apical tufts 0.5–1 mm, abaxial surfaces white, long hairs common to abundant, cottony-crisped hairs dense, short hairs and glands absent or obscured, adaxial grayish green, long hairs common to abundant, 0.5–1.5 mm, ± stiff, rarely soft, short and/or crisped hairs sparse to common, cottony hairs absent, glands sparse to common. |
Cauline leaves | (0–)1–2. |
1–2. |
Inflorescences | 3–6(–10)-flowered, loosely cymose, sometimes racemiform. |
(1–)3–10-flowered, congested or ± elongating in fruit, branch angle 20–35°. |
Pedicels | (0.5–)1–2(–4.5) cm, ± recurved in fruit. |
0.2–0.5 cm, proximal to 1.2 cm. |
Flowers | epicalyx bractlets ± elliptic, 2–4(–6) × 1–2(–2.5) mm; hypanthium 3–6 mm diam.; sepals 4–6(–8) mm, apex acute; petals 4–8(–10) × 3–7(–9) mm; filaments 2–3.5 mm, anthers 0.7–1 mm; carpels 10–30, styles (1.5–)2–3 mm. |
epicalyx bractlets narrowly elliptic, 1.5–3 × 0.5–1 mm; hypanthium 3–4 mm diam.; sepals 2.5–4 mm, apex ± acute, glands abundant, usually not obscured; petals yellow, ± overlapping, 3.5–5 × 3–4 mm, slightly longer than sepals; filaments 1 mm, anthers 0.3–0.7 mm; carpels 20–40, styles 1 mm. |
Achenes | 1.5–2 mm, smooth, often ± carunculate. |
1 mm. |
Potentilla millefolia |
Potentilla modesta |
|
Phenology | Flowering spring–summer. | Flowering summer. |
Habitat | Vernally to permanently wet meadows, moist openings in conifer forests and sagebrush, alkaline flats | Alpine tundra, fellfields, talus slopes, cliffs, usually on limestone |
Elevation | 700–2200 m (2300–7200 ft) | 2500–3900 m (8200–12800 ft) |
Distribution |
CA; NV; OR
|
CO; ID; MT; UT; WY |
Discussion | Potentilla millefolia occurs from central Oregon to the east side of the Sierra Nevada of California, with a disjunct occurrence on the alkaline flats of Reese River Valley, Nevada. Significant variation occurs in vestiture type, leaflet dissection, and flower size, but with minimal geographic correlation. The most distinctive variant, represented by the type of P. klamathensis, has relatively long, slender, spreading, pustule-based hairs, often intermixed with shorter hairs. This vestiture type does not appear to be correlated with any other characters or geographic distribution and may vary within a population. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Potentilla modesta is the dominant component of what has previously been called P. rubricaulis in the Intermountain Region (for example, N. H. Holmgren 1997b, including illustration). This and other species of alpine Potentilla often grow in mixed populations, which contributes to confusion and mistaken identifications. In general, P. modesta has a more compact inflorescence and more consistently palmate basal leaves than sympatric members of the section. Intergrades with P. jepsonii (sect. Pensylvanicae) can also be problematic, though leaves of the latter are generally subpinnate and abaxially grayish rather than white. The epithet modesta is misapplied in the combination Potentilla concinna var. modesta, which S. L. Welsh et al. (1993) used for plants mostly placed here in P. concinna var. divisa (sect. Concinnae). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 9, p. 174. | FNA vol. 9, p. 209. |
Parent taxa | Rosaceae > subfam. Rosoideae > tribe Potentilleae > Potentilla > sect. Multijugae | Rosaceae > subfam. Rosoideae > tribe Potentilleae > Potentilla > sect. Rubricaules |
Sibling taxa | ||
Synonyms | P. klamathensis, P. millefolia var. klamathensis, P. plattensis var. klamathensis, P. plattensis var. millefolia | P. concinna var. modesta |
Name authority | Rydberg: Bull. Torrey Bot. Club 23: 433, plate 277, figs. 1–5. (1896) | Rydberg: in N. L. Britton et al., N. Amer. Fl. 22: 331. (1908) |
Web links |