Potentilla millefolia |
Potentilla lasiodonta |
|
---|---|---|
cut-leaf cinquefoil, feather cinquefoil, feather or many-leaf or Klamath cinquefoil, many leaf cinquefoil |
Sandhills cinquefoil |
|
Habit | Plants ± rosetted; taproots fleshy-thickened. | |
Stems | usually prostrate, sometimes ± decumbent, 0.4–2(–3) dm, lengths 1–2 times basal leaves. |
± erect, 2–4 dm. |
Basal leaves | pinnate with distal leaflets ± confluent, 2–15(–20) × 1–3 cm; petiole 0.5–2(–3) cm, straight hairs sparse to abundant, appressed to spreading, 0.5–1.5 mm, stiff, cottony hairs absent, glands absent or sparse; primary lateral leaflets (3–)5–13 per side, on distal 2/3–3/4+ of leaf axis, separate to overlapping, largest ones cuneate to flabellate, 0.5–1.5(–2) × 0.5–2 cm, distal 2/3 to whole margin palmately or unevenly, rarely pinnately, incised 2/3 to completely to midvein, ultimate teeth or segments (1–)2–10, linear to broadly oblanceolate, 2–10 × (0.5–)1–2 mm, apical tufts to 1 mm, surfaces green to grayish green, not glaucous, straight hairs sparse to abundant, appressed to spreading, 0.5–1.5(–2) mm, stiff, cottony hairs absent, glands sparse to common. |
pinnate, 12–20(–30) cm; petiole 5–10(–15) cm, long hairs ± abundant, spreading to ascending, 0.5–1 mm, weak to ± stiff, short hairs abundant to dense, cottony or crisped hairs absent, glands sparse to abundant, often obscured; leaflets 4–6(–9) per side, on distal (1/3–)1/2–3/5 of leaf axis, slightly overlapping, terminal ones oblanceolate to narrowly elliptic, 3–6 × 0.8–1.2(–1.5) cm, margins strongly revolute, incised ± 1/2 to midvein, undivided medial blade 5–9 mm wide, teeth 8–12 per side, narrowly triangular to lanceolate, surfaces ± dissimilar, abaxial grayish, long hairs ± sparse (or not differentiated from short hairs), 0.5–1.5 mm, weak, short hairs abundant to dense, cottony or crisped hairs absent, glands sparse or obscured, adaxial ± green, straight hairs (long and short not differentiated) abundant, spreading to ascending, 0.2–0.5 mm, cottony and crisped hairs absent, glands sparse to abundant, often obscured. |
Cauline leaves | (0–)1–2. |
(1–)2–4. |
Inflorescences | 3–6(–10)-flowered, loosely cymose, sometimes racemiform. |
10–40-flowered, congested or elongating in fruit. |
Pedicels | (0.5–)1–2(–4.5) cm, ± recurved in fruit. |
± 0.1 cm (proximal to 1.2 cm). |
Flowers | epicalyx bractlets ± elliptic, 2–4(–6) × 1–2(–2.5) mm; hypanthium 3–6 mm diam.; sepals 4–6(–8) mm, apex acute; petals 4–8(–10) × 3–7(–9) mm; filaments 2–3.5 mm, anthers 0.7–1 mm; carpels 10–30, styles (1.5–)2–3 mm. |
epicalyx bractlets narrowly ovate-acuminate, (3–)4–8(–10) × 2–3 mm, lengths 1–2 times sepals, margins ± revolute; hypanthium (4–)5–10 mm diam.; sepals 3–5 mm, apex acute to obtuse, abaxial surfaces: venation moderate, glands ± abundant, obscured to evident; petals pale yellow, 3–5 × 2–4 mm, lengths ± equal to or shorter than sepals; filaments 1–1.5 mm, anthers 0.5–0.8 mm; carpels 80–90, styles papillate-swollen in proximal 3/4+, 1–1.2 mm. |
Achenes | 1.5–2 mm, smooth, often ± carunculate. |
± 1 mm, ± rugose. |
2n | = 14. |
|
Potentilla millefolia |
Potentilla lasiodonta |
|
Phenology | Flowering spring–summer. | Flowering summer. |
Habitat | Vernally to permanently wet meadows, moist openings in conifer forests and sagebrush, alkaline flats | Sandy sites in prairies |
Elevation | 700–2200 m (2300–7200 ft) | 300–1100 m (1000–3600 ft) |
Distribution |
CA; NV; OR
|
MN; ND; AB; MB; SK |
Discussion | Potentilla millefolia occurs from central Oregon to the east side of the Sierra Nevada of California, with a disjunct occurrence on the alkaline flats of Reese River Valley, Nevada. Significant variation occurs in vestiture type, leaflet dissection, and flower size, but with minimal geographic correlation. The most distinctive variant, represented by the type of P. klamathensis, has relatively long, slender, spreading, pustule-based hairs, often intermixed with shorter hairs. This vestiture type does not appear to be correlated with any other characters or geographic distribution and may vary within a population. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Potentilla lasiodonta is a diploid relative of tetraploid P. pensylvanica, sharing similar velvety vestiture of dense short hairs, pinnate leaves, and epicalyx bractlets with revolute margins. The species differs in having larger epicalyx bractlets and less deeply incised leaflets. Plant height and leaf size are at the upper range of P. pensylvanica, and populations are evidently restricted to sandy substrates. Occurrence in Manitoba is based on B. L. Kohli and J. G. Packer (1976); no vouchers are in WIN. At least one collection from southeastern British Columbia (Brown 779, MO) approaches P. lasiodonta, but it is insufficient by itself to serve as a provincial record. The fundamental ploidy and epicalyx distinctions of this species were established by B. L. Kohli and J. G. Packer (1976), who proposed the name Potentilla finitima. As noted by J. Soják (1994), the type of P. lasiodonta is the same entity. Potentilla atrovirens Rydberg and P. pensylvanica var. arida B. Boivin have sometimes been applied to this taxon; the types of both names fall within the circumscription of P. pensylvanica in the narrow sense (Packer, pers. comm.). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 9, p. 174. | FNA vol. 9, p. 214. |
Parent taxa | Rosaceae > subfam. Rosoideae > tribe Potentilleae > Potentilla > sect. Multijugae | Rosaceae > subfam. Rosoideae > tribe Potentilleae > Potentilla > sect. Pensylvanicae |
Sibling taxa | ||
Synonyms | P. klamathensis, P. millefolia var. klamathensis, P. plattensis var. klamathensis, P. plattensis var. millefolia | P. finitima |
Name authority | Rydberg: Bull. Torrey Bot. Club 23: 433, plate 277, figs. 1–5. (1896) | Rydberg: in N. L. Britton et al., N. Amer. Fl. 22: 351. (1908) |
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