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cut-leaf cinquefoil, feather cinquefoil, feather or many-leaf or Klamath cinquefoil, many leaf cinquefoil

Jepson's cinquefoil, Jepson's potentilla

Habit Plants ± rosetted; taproots fleshy-thickened.
Stems

usually prostrate, sometimes ± decumbent, 0.4–2(–3) dm, lengths 1–2 times basal leaves.

decumbent to nearly erect, (0.2–)0.5–2.6 dm.

Basal leaves

pinnate with distal leaflets ± confluent, 2–15(–20) × 1–3 cm;

petiole 0.5–2(–3) cm, straight hairs sparse to abundant, appressed to spreading, 0.5–1.5 mm, stiff, cottony hairs absent, glands absent or sparse;

primary lateral leaflets (3–)5–13 per side, on distal 2/3–3/4+ of leaf axis, separate to overlapping, largest ones cuneate to flabellate, 0.5–1.5(–2) × 0.5–2 cm, distal 2/3 to whole margin palmately or unevenly, rarely pinnately, incised 2/3 to completely to midvein, ultimate teeth or segments (1–)2–10, linear to broadly oblanceolate, 2–10 × (0.5–)1–2 mm, apical tufts to 1 mm, surfaces green to grayish green, not glaucous, straight hairs sparse to abundant, appressed to spreading, 0.5–1.5(–2) mm, stiff, cottony hairs absent, glands sparse to common.

subpinnate to subpalmate, 1.5–10(–17) cm;

petiole 0.5–4(–8) cm, long hairs ± abundant, spreading to loosely appressed, 0.5–2 mm, weak to stiff, short hairs absent or sparse, cottony or crisped hairs absent, glands sparse to abundant;

leaflets (1–)2–3 per side, on distal 1/6–1/3(–1/2) of leaf axis, separate to slightly overlapping, terminal ones oblanceolate-elliptic to narrowly obovate, (0.5–)1–2.5(–3.5) × 0.4–1(–1.3) cm, margins strongly revolute, incised 3/4+ to midvein, undivided medial blade 1.5–3(–5) mm wide, teeth (2–)3–5(–6) per side, ± linear-elliptic to rarely ovate or oblanceolate, surfaces ± dissimilar, abaxial ± grayish, rarely white, long hairs ± abundant especially on veins, 1 mm, weak, short hairs absent or sparse, cottony hairs absent, crisped hairs sparse to abundant, glands ± abundant, sometimes obscured, adaxial green to gray-green, long hairs sparse to abundant, loosely appressed, 0.5–1(–2) mm, short hairs absent or sparse, cottony and crisped hairs absent, glands ± abundant.

Cauline leaves

(0–)1–2.

1–2.

Inflorescences

3–6(–10)-flowered, loosely cymose, sometimes racemiform.

(1–)3–12(–20)-flowered, congested, scarcely opening in fruit.

Pedicels

(0.5–)1–2(–4.5) cm, ± recurved in fruit.

0.2–0.5 cm (proximal to 1.2 cm).

Flowers

epicalyx bractlets ± elliptic, 2–4(–6) × 1–2(–2.5) mm;

hypanthium 3–6 mm diam.;

sepals 4–6(–8) mm, apex acute;

petals 4–8(–10) × 3–7(–9) mm;

filaments 2–3.5 mm, anthers 0.7–1 mm;

carpels 10–30, styles (1.5–)2–3 mm.

epicalyx bractlets elliptic to linear, 1.5–3 × (0.3–)0.5–1.2 mm, lengths 1/2–3/4 times sepals, margins flat;

hypanthium 2.5–5 mm diam.;

sepals 2–5 mm, apex usually ± acute to obtuse, rarely acuminate, abaxial surfaces: venation faint, glands abundant, evident;

petals pale yellow, 2–5 × 2–4 mm, lengths ± equal to sepals;

filaments 0.5–2 mm, anthers 0.3–0.7 mm;

carpels 20–60, styles papillate-swollen in proximal 1/2–3/4+, 0.8–1.2 mm.

Achenes

1.5–2 mm, smooth, often ± carunculate.

1 mm, smooth to faintly rugose.

Potentilla millefolia

Potentilla jepsonii

Phenology Flowering spring–summer. Flowering summer.
Habitat Vernally to permanently wet meadows, moist openings in conifer forests and sagebrush, alkaline flats Seasonally moist sites in alpine meadows, ridges, fellfields, rock ledges, talus slopes
Elevation 700–2200 m (2300–7200 ft) 2500–3700 m (8200–12100 ft)
Distribution
from FNA
CA; NV; OR
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; CA; CO; ID; MT; NV; OR; UT; WA; AB
[BONAP county map]
Discussion

Potentilla millefolia occurs from central Oregon to the east side of the Sierra Nevada of California, with a disjunct occurrence on the alkaline flats of Reese River Valley, Nevada. Significant variation occurs in vestiture type, leaflet dissection, and flower size, but with minimal geographic correlation. The most distinctive variant, represented by the type of P. klamathensis, has relatively long, slender, spreading, pustule-based hairs, often intermixed with shorter hairs. This vestiture type does not appear to be correlated with any other characters or geographic distribution and may vary within a population.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Potentilla jepsonii (a replacement name for P. pensylvanica var. ovium) is widespread at higher elevations in the Intermountain Region, extending into the Sierra Nevada in California and the Rocky Mountains in Idaho and Wyoming. Disjunct occurrences are known on the north rim of the Grand Canyon and San Francisco Peaks, Arizona, and Steens Mountain, Oregon. Collections have most commonly been included in P. pensylvanica (for example, B. Ertter 1993; N. H. Holmgren 1997b); they differ in leaf division, vestiture, and epicalyx bractlets. J. Soják considered var. ovium to be a variety of P. litoralis, which has comparable leaves and vestiture; P. jepsonii lacks the enlarged, revolute epicalyx bractlets that distinguish P. litoralis and is for the most part well separated geographically and elevationally. The exception is the montane phase of P. litoralis, which, at the current stage of knowledge, is somewhat arbitrarily delimited from P. jepsonii in the northern Rocky Mountains; collections from Waterton Lakes National Park in Alberta and the Okanogan Mountains in Washington are tentatively placed here in P. jepsonii.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 9, p. 174. FNA vol. 9, p. 216.
Parent taxa Rosaceae > subfam. Rosoideae > tribe Potentilleae > Potentilla > sect. Multijugae Rosaceae > subfam. Rosoideae > tribe Potentilleae > Potentilla > sect. Pensylvanicae
Sibling taxa
P. albiflora, P. ambigens, P. anachoretica, P. angelliae, P. anglica, P. anserina, P. arenosa, P. argentea, P. arizonica, P. basaltica, P. bicrenata, P. biennis, P. biflora, P. bimundorum, P. bipinnatifida, P. brevifolia, P. breweri, P. bruceae, P. brunnescens, P. canadensis, P. concinna, P. cottamii, P. crantzii, P. crebridens, P. crinita, P. cristae, P. demotica, P. drummondii, P. effusa, P. elegans, P. erecta, P. flabellifolia, P. fragiformis, P. furcata, P. glaucophylla, P. gracilis, P. grayi, P. hickmanii, P. hippiana, P. holmgrenii, P. hookeriana, P. hyparctica, P. inclinata, P. intermedia, P. jepsonii, P. johnstonii, P. lasiodonta, P. litoralis, P. macounii, P. modesta, P. morefieldii, P. multijuga, P. multisecta, P. nana, P. newberryi, P. nivea, P. norvegica, P. ovina, P. paucijuga, P. pedersenii, P. pensylvanica, P. plattensis, P. pseudosericea, P. pulchella, P. pulcherrima, P. recta, P. reptans, P. rhyolitica, P. rimicola, P. rivalis, P. robbinsiana, P. rubella, P. rubricaulis, P. sanguinea, P. saximontana, P. sierrae-blancae, P. simplex, P. sterilis, P. stipularis, P. subgorodkovii, P. subjuga, P. subvahliana, P. subviscosa, P. supina, P. thurberi, P. thuringiaca, P. tikhomirovii, P. townsendii, P. uliginosa, P. uschakovii, P. vahliana, P. verna, P. versicolor, P. villosa, P. villosula, P. vulcanicola, P. wheeleri
P. albiflora, P. ambigens, P. anachoretica, P. angelliae, P. anglica, P. anserina, P. arenosa, P. argentea, P. arizonica, P. basaltica, P. bicrenata, P. biennis, P. biflora, P. bimundorum, P. bipinnatifida, P. brevifolia, P. breweri, P. bruceae, P. brunnescens, P. canadensis, P. concinna, P. cottamii, P. crantzii, P. crebridens, P. crinita, P. cristae, P. demotica, P. drummondii, P. effusa, P. elegans, P. erecta, P. flabellifolia, P. fragiformis, P. furcata, P. glaucophylla, P. gracilis, P. grayi, P. hickmanii, P. hippiana, P. holmgrenii, P. hookeriana, P. hyparctica, P. inclinata, P. intermedia, P. johnstonii, P. lasiodonta, P. litoralis, P. macounii, P. millefolia, P. modesta, P. morefieldii, P. multijuga, P. multisecta, P. nana, P. newberryi, P. nivea, P. norvegica, P. ovina, P. paucijuga, P. pedersenii, P. pensylvanica, P. plattensis, P. pseudosericea, P. pulchella, P. pulcherrima, P. recta, P. reptans, P. rhyolitica, P. rimicola, P. rivalis, P. robbinsiana, P. rubella, P. rubricaulis, P. sanguinea, P. saximontana, P. sierrae-blancae, P. simplex, P. sterilis, P. stipularis, P. subgorodkovii, P. subjuga, P. subvahliana, P. subviscosa, P. supina, P. thurberi, P. thuringiaca, P. tikhomirovii, P. townsendii, P. uliginosa, P. uschakovii, P. vahliana, P. verna, P. versicolor, P. villosa, P. villosula, P. vulcanicola, P. wheeleri
Synonyms P. klamathensis, P. millefolia var. klamathensis, P. plattensis var. klamathensis, P. plattensis var. millefolia P. pensylvanica var. ovium, P. litoralis var. ovium
Name authority Rydberg: Bull. Torrey Bot. Club 23: 433, plate 277, figs. 1–5. (1896) Ertter: J. Bot. Res. Inst. Texas 2: 202. (2008)
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