Potentilla millefolia |
Potentilla holmgrenii |
|
---|---|---|
cut-leaf cinquefoil, feather cinquefoil, feather or many-leaf or Klamath cinquefoil, many leaf cinquefoil |
Holmgren's cinquefoil |
|
Habit | Plants ± rosetted; taproots fleshy-thickened. | Plants densely tufted to cushion-forming. |
Caudex branches | stout, columnar, at least partly sheathed with marcescent whole leaves. |
|
Stems | usually prostrate, sometimes ± decumbent, 0.4–2(–3) dm, lengths 1–2 times basal leaves. |
ascending, 0.2–0.8 dm, lengths 1.5–2.5 times basal leaves. |
Basal leaves | pinnate with distal leaflets ± confluent, 2–15(–20) × 1–3 cm; petiole 0.5–2(–3) cm, straight hairs sparse to abundant, appressed to spreading, 0.5–1.5 mm, stiff, cottony hairs absent, glands absent or sparse; primary lateral leaflets (3–)5–13 per side, on distal 2/3–3/4+ of leaf axis, separate to overlapping, largest ones cuneate to flabellate, 0.5–1.5(–2) × 0.5–2 cm, distal 2/3 to whole margin palmately or unevenly, rarely pinnately, incised 2/3 to completely to midvein, ultimate teeth or segments (1–)2–10, linear to broadly oblanceolate, 2–10 × (0.5–)1–2 mm, apical tufts to 1 mm, surfaces green to grayish green, not glaucous, straight hairs sparse to abundant, appressed to spreading, 0.5–1.5(–2) mm, stiff, cottony hairs absent, glands sparse to common. |
1.5–4 cm; petiole 0.5–2 cm, long hairs absent or sparse to common (less so than cottony hairs), ± appressed, 1–2 mm, soft, smooth, cottony hairs dense, other hairs and glands absent, sparse, or obscured; leaflets touching to overlapping, central broadly elliptic to obovate, 0.7–1.1 × 0.5–0.6 cm, sessile or short-petiolulate, petiolule to 1 mm, base cuneate, margins revolute, distal 1/2–3/4 incised ± 1/2 to midvein, teeth 2–4 per side, approximate, surfaces ± dissimilar, abaxial white to grayish white, long hairs 1 mm, cottony-crisped hairs dense, adaxial grayish green to grayish white, long and short-crisped hairs abundant to dense. |
Cauline leaves | (0–)1–2. |
0–1. |
Inflorescences | 3–6(–10)-flowered, loosely cymose, sometimes racemiform. |
1–2(–3)-flowered. |
Pedicels | (0.5–)1–2(–4.5) cm, ± recurved in fruit. |
0.4–0.7 cm in flower, to 2 cm in fruit. |
Flowers | epicalyx bractlets ± elliptic, 2–4(–6) × 1–2(–2.5) mm; hypanthium 3–6 mm diam.; sepals 4–6(–8) mm, apex acute; petals 4–8(–10) × 3–7(–9) mm; filaments 2–3.5 mm, anthers 0.7–1 mm; carpels 10–30, styles (1.5–)2–3 mm. |
epicalyx bractlets lanceolate to narrowly elliptic, 2–2.2 × 0.6–0.9 mm, 1/4–1/2 as wide as sepals, margins flat, red glands absent; hypanthium 1.8–2.2 mm diam.; sepals 3–3.5 mm, apex subacute; petals 4–5 × 2.5–4 mm, longer than sepals; filaments 1–1.5 mm, anthers 0.5 mm; carpels 30–40, apical hairs absent, styles narrowly columnar, not or ± papillate-swollen in proximal 1/5 or less, 1.2–1.4 mm. |
Achenes | 1.5–2 mm, smooth, often ± carunculate. |
1.1–1.4 mm. |
Potentilla millefolia |
Potentilla holmgrenii |
|
Phenology | Flowering spring–summer. | Flowering summer. |
Habitat | Vernally to permanently wet meadows, moist openings in conifer forests and sagebrush, alkaline flats | Wind-swept ridges, fellfields, rocky slopes |
Elevation | 700–2200 m (2300–7200 ft) | 3300–4500 m (10800–14800 ft) |
Distribution |
CA; NV; OR
|
NV; UT |
Discussion | Potentilla millefolia occurs from central Oregon to the east side of the Sierra Nevada of California, with a disjunct occurrence on the alkaline flats of Reese River Valley, Nevada. Significant variation occurs in vestiture type, leaflet dissection, and flower size, but with minimal geographic correlation. The most distinctive variant, represented by the type of P. klamathensis, has relatively long, slender, spreading, pustule-based hairs, often intermixed with shorter hairs. This vestiture type does not appear to be correlated with any other characters or geographic distribution and may vary within a population. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. Potentilla holmgrenii is known only from the Schell Creek and Snake ranges, Nevada, and Deep Creek Range, Utah. In addition to characteristics given in the key, lateral leaflets and leaflet teeth of P. holmgrenii are directed more forward than in P. nivea, leaflets are thicker and more densely hairy, and the terminal tooth is often orbiculate and frequently overlapped by the adjacent pair of lateral teeth. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 9, p. 174. | FNA vol. 9, p. 199. |
Parent taxa | Rosaceae > subfam. Rosoideae > tribe Potentilleae > Potentilla > sect. Multijugae | Rosaceae > subfam. Rosoideae > tribe Potentilleae > Potentilla > sect. Niveae |
Sibling taxa | ||
Synonyms | P. klamathensis, P. millefolia var. klamathensis, P. plattensis var. klamathensis, P. plattensis var. millefolia | |
Name authority | Rydberg: Bull. Torrey Bot. Club 23: 433, plate 277, figs. 1–5. (1896) | D. F. Murray & Elven: J. Bot. Res. Inst. Texas 1: 811, figs. 1, 2. (2007) |
Web links |