Potentilla millefolia |
Potentilla effusa |
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cut-leaf cinquefoil, feather cinquefoil, feather or many-leaf or Klamath cinquefoil, many leaf cinquefoil |
branch cinquefoil |
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Habit | Plants ± rosetted; taproots fleshy-thickened. | |||||
Stems | usually prostrate, sometimes ± decumbent, 0.4–2(–3) dm, lengths 1–2 times basal leaves. |
(0.5–)1.5–4(–5.5) dm, lengths 2–3(–4) times basal leaves. |
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Basal leaves | pinnate with distal leaflets ± confluent, 2–15(–20) × 1–3 cm; petiole 0.5–2(–3) cm, straight hairs sparse to abundant, appressed to spreading, 0.5–1.5 mm, stiff, cottony hairs absent, glands absent or sparse; primary lateral leaflets (3–)5–13 per side, on distal 2/3–3/4+ of leaf axis, separate to overlapping, largest ones cuneate to flabellate, 0.5–1.5(–2) × 0.5–2 cm, distal 2/3 to whole margin palmately or unevenly, rarely pinnately, incised 2/3 to completely to midvein, ultimate teeth or segments (1–)2–10, linear to broadly oblanceolate, 2–10 × (0.5–)1–2 mm, apical tufts to 1 mm, surfaces green to grayish green, not glaucous, straight hairs sparse to abundant, appressed to spreading, 0.5–1.5(–2) mm, stiff, cottony hairs absent, glands sparse to common. |
pinnate, (3–)5–15(–25) cm; petiole 0.5–5(–8) cm, long hairs absent to abundant, ± appressed, 1–3 mm, stiff to weak, short and crisped hairs absent or obscured, cottony hairs abundant (at least on first-formed leaves), glands sparse or obscured; leaflets not conduplicate, lateral ones evenly to unevenly paired, 2–5(–7) per side (secondary leaflets sometimes interspersed) on distal 1/3–1/2 of leaf axis, distal pairs usually not decurrent or confluent with terminal leaflet, larger leaflets oblanceolate to narrowly obovate, 1–2.5(–3) × 0.3–1.3(–1.8) cm, distal (1/2–)2/3–3/4 (rarely more) of margin incised ± 1/2 to midvein, teeth (1–)2–9 per side, 1–4 mm, surfaces similar, gray to white or ± green, abaxial: long hairs sparse to common (at least on veins), 0.5–1.5 mm, weak to stiff, short or crisped hairs absent or sparse, sometimes obscured, cottony hairs absent or sparse to dense, glands absent, sparse, or obscured, adaxial: long hairs absent or sparse to common, short or crisped hairs absent or sparse, cottony hairs absent or sparse to abundant, glands absent or sparse. |
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Cauline leaves | (0–)1–2. |
2–6+. |
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Inflorescences | 3–6(–10)-flowered, loosely cymose, sometimes racemiform. |
7–30-flowered. |
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Pedicels | (0.5–)1–2(–4.5) cm, ± recurved in fruit. |
0.3–3 cm. |
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Flowers | epicalyx bractlets ± elliptic, 2–4(–6) × 1–2(–2.5) mm; hypanthium 3–6 mm diam.; sepals 4–6(–8) mm, apex acute; petals 4–8(–10) × 3–7(–9) mm; filaments 2–3.5 mm, anthers 0.7–1 mm; carpels 10–30, styles (1.5–)2–3 mm. |
epicalyx bractlets linear to narrowly lanceolate, 1–2.5(–3) × 0.3–1 mm, 1/2–2/3 as long as sepals, abaxial vestiture often much sparser than sepals, often glabrate or glabrescent distally, straight hairs absent or sparse, cottony hairs usually abundant (at least proximally), sometimes absent or nearly so (var. rupincola); hypanthium 2–6 mm diam.; sepals 3–6(–7) mm, apex long acuminate; petals 4–6.5 × 4–6 mm; filaments 1–3.5 mm, anthers 0.5–1 mm; carpels 3–15, styles 1.5–2.1 mm. |
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Achenes | 1.5–2 mm, smooth, often ± carunculate. |
1.8–2.1 mm, smooth. |
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Potentilla millefolia |
Potentilla effusa |
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Phenology | Flowering spring–summer. | |||||
Habitat | Vernally to permanently wet meadows, moist openings in conifer forests and sagebrush, alkaline flats | |||||
Elevation | 700–2200 m (2300–7200 ft) | |||||
Distribution |
CA; NV; OR
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CO; ID; MN; MT; ND; NE; NM; SD; UT; WY; AB; MB; SK |
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Discussion | Potentilla millefolia occurs from central Oregon to the east side of the Sierra Nevada of California, with a disjunct occurrence on the alkaline flats of Reese River Valley, Nevada. Significant variation occurs in vestiture type, leaflet dissection, and flower size, but with minimal geographic correlation. The most distinctive variant, represented by the type of P. klamathensis, has relatively long, slender, spreading, pustule-based hairs, often intermixed with shorter hairs. This vestiture type does not appear to be correlated with any other characters or geographic distribution and may vary within a population. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Varieties 2 (2 in the flora). In the southern Rocky Mountains, Potentilla effusa is relatively distinct from P. hippiana both morphologically and geographically, with P. effusa mostly east of the Continental Divide and P. hippiana mostly west. In general, P. hippiana differs from P. effusa in having leaflets more evenly paired, more tightly serrate with teeth often occurring on the whole margin, and more bicolored with abaxial vestiture of crisped rather than cottony hairs. Stems of P. effusa have more cauline leaves on average than P. hippiana, and vestiture of epicalyx bractlets of P. effusa (as here circumscribed) is never sericeous. The distinction between the two species is marred by intermediate populations throughout their shared range in the northern Rocky Mountains and Great Plains. Most regional floras focused on these areas have accordingly included Potentilla effusa within P. hippiana, sometimes without infraspecific distinction. An alternate solution has been the recognition of P. hippiana subsp. effusa, with the subspecies further subdivided into the two varieties recognized here. The varieties themselves intergrade; P. effusa var. ×coloradensis (Rydberg) Th. Wolf is available for the hybrid. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 9, p. 174. | FNA vol. 9, p. 162. | ||||
Parent taxa | Rosaceae > subfam. Rosoideae > tribe Potentilleae > Potentilla > sect. Multijugae | Rosaceae > subfam. Rosoideae > tribe Potentilleae > Potentilla > sect. Leucophyllae | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | P. klamathensis, P. millefolia var. klamathensis, P. plattensis var. klamathensis, P. plattensis var. millefolia | P. hippiana subsp. effusa, P. hippiana var. effusa | ||||
Name authority | Rydberg: Bull. Torrey Bot. Club 23: 433, plate 277, figs. 1–5. (1896) | Douglas ex Lehmann: Nov. Stirp. Pug. 2: 8. (1830) | ||||
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