Potentilla millefolia |
Potentilla drummondii |
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cut-leaf cinquefoil, feather cinquefoil, feather or many-leaf or Klamath cinquefoil, many leaf cinquefoil |
Drummond's cinquefoil |
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Habit | Plants ± rosetted; taproots fleshy-thickened. | |
Glands | usually absent or inconspicuous, rarely conspicuous, uncolored. |
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Stems | usually prostrate, sometimes ± decumbent, 0.4–2(–3) dm, lengths 1–2 times basal leaves. |
decumbent to nearly erect, 1.5–4.5(–6) dm. |
Basal leaves | pinnate with distal leaflets ± confluent, 2–15(–20) × 1–3 cm; petiole 0.5–2(–3) cm, straight hairs sparse to abundant, appressed to spreading, 0.5–1.5 mm, stiff, cottony hairs absent, glands absent or sparse; primary lateral leaflets (3–)5–13 per side, on distal 2/3–3/4+ of leaf axis, separate to overlapping, largest ones cuneate to flabellate, 0.5–1.5(–2) × 0.5–2 cm, distal 2/3 to whole margin palmately or unevenly, rarely pinnately, incised 2/3 to completely to midvein, ultimate teeth or segments (1–)2–10, linear to broadly oblanceolate, 2–10 × (0.5–)1–2 mm, apical tufts to 1 mm, surfaces green to grayish green, not glaucous, straight hairs sparse to abundant, appressed to spreading, 0.5–1.5(–2) mm, stiff, cottony hairs absent, glands sparse to common. |
sometimes 2-ranked, subpinnate to pinnate (proximal leaflets often doubled, distal leaflets confluent and/or decurrent), (4–)10–25 cm; petiole 1–10(–15) cm, long hairs absent or sparse to abundant, appressed, 1–1.5 mm, usually stiff, short, crisped, and cottony hairs usually absent, glands absent or (regionally) abundant; leaflets 5–9, on distal 1/10–1/2(–3/4) of leaf axis, separate to ± overlapping, largest ones obovate to cuneate, (1–)2–5 × (0.7–)1–3(–3.5) cm, margins flat, distal 1/2–3/4 unevenly, sometimes evenly, incised ± 1/2 to midvein (often with additional incisions nearly to midvein), undivided medial blade 3–15 mm wide, teeth 3–7 per side (sometimes secondarily toothed), linear to lanceolate, 3–10 mm, surfaces similar, green, not glaucous, long hairs nearly absent or sparse to common (sometimes restricted to abaxial veins), short, crisped, and cottony hairs usually absent, glands usually absent, sometimes regionally abundant. |
Cauline leaves | (0–)1–2. |
1–3. |
Inflorescences | 3–6(–10)-flowered, loosely cymose, sometimes racemiform. |
3–15-flowered. |
Pedicels | (0.5–)1–2(–4.5) cm, ± recurved in fruit. |
1–3(–6.5) cm. |
Flowers | epicalyx bractlets ± elliptic, 2–4(–6) × 1–2(–2.5) mm; hypanthium 3–6 mm diam.; sepals 4–6(–8) mm, apex acute; petals 4–8(–10) × 3–7(–9) mm; filaments 2–3.5 mm, anthers 0.7–1 mm; carpels 10–30, styles (1.5–)2–3 mm. |
epicalyx bractlets lanceolate to narrowly elliptic, 3–5 × 1–1.6 mm, hairs usually sparse, appressed to ascending, glands usually absent, sometimes regionally sparse to common; hypanthium (3–)4–6(–7) mm diam.; sepals 5–10 mm, apex acute to acuminate; petals (6–)7–12 × 5–10 mm; filaments 2–3.5 mm, anthers 0.7–1 mm; carpels (20–)25–40, styles filiform above papillate-swollen base, 2–3 mm. |
Achenes | 1.5–2 mm, smooth, often ± carunculate. |
1.5–2 mm. |
2n | = 64, 92, 96, 98–108. |
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Potentilla millefolia |
Potentilla drummondii |
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Phenology | Flowering spring–summer. | Flowering summer. |
Habitat | Vernally to permanently wet meadows, moist openings in conifer forests and sagebrush, alkaline flats | Moist to dry meadows and adjacent slopes, in conifer woodlands, alpine tundra communities |
Elevation | 700–2200 m (2300–7200 ft) | 300–3000 m (1000–9800 ft) |
Distribution |
CA; NV; OR
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AK; CA; MT; OR; WA; AB; BC
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Discussion | Potentilla millefolia occurs from central Oregon to the east side of the Sierra Nevada of California, with a disjunct occurrence on the alkaline flats of Reese River Valley, Nevada. Significant variation occurs in vestiture type, leaflet dissection, and flower size, but with minimal geographic correlation. The most distinctive variant, represented by the type of P. klamathensis, has relatively long, slender, spreading, pustule-based hairs, often intermixed with shorter hairs. This vestiture type does not appear to be correlated with any other characters or geographic distribution and may vary within a population. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
The present circumscription of Potentilla drummondii differs from that of B. Ertter (1992, 1993), which encompassed P. breweri and P. bruceae as varieties (see 28. P. bruceae discussion for details). The description above focuses on large, eglandular, subpinnate plants that occur in mountains from the central Sierra Nevada and northern Coast Ranges of California to the Olympic Peninsula of Washington, east to southern Alberta. At least some collections previously identified as P. glaucophylla (as P. diversifolia Lehmann) from the Kenai Peninsula in Alaska can be readily accommodated within this concept, as can collections from the Cabinet Mountains in Montana. The distinction between P. drummondii and subpalmate P. glaucophylla can be vague; in general, the proximalmost leaflets of P. glaucophylla are relatively small and often entire; in P. drummondii proximalmost leaflets are similar in size and dissection to other leaflets. Pinnate-leaved regional variants merit further attention; in particular the smaller form described as Potentilla cascadensis, and glandular plants in the northern Coast Ranges of California. Relatively small plants forming uniform populations in the southern Sierra Nevada may represent stabilized hybrids with P. breweri. The inclusion of Potentilla bruceae and P. drummondii in sect. Graciles differs from that of J. Clausen et al. (1940) and B. C. Johnston (1980, 1985), who allied both species with P. breweri in sect. Multijugae. Although subpalmate to subpinnate leaves are anomalous in primarily palmate sect. Graciles, doubled proximal leaflets and an indistinct transition between the rachis and axis of the pinnatisect terminal leaflet in these two species suggest that this state is derived from a palmate progenitor. Habit and petioles also are more compatible with sect. Graciles than sect. Multijugae. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 9, p. 174. | FNA vol. 9, p. 159. |
Parent taxa | Rosaceae > subfam. Rosoideae > tribe Potentilleae > Potentilla > sect. Multijugae | Rosaceae > subfam. Rosoideae > tribe Potentilleae > Potentilla > sect. Graciles |
Sibling taxa | ||
Synonyms | P. klamathensis, P. millefolia var. klamathensis, P. plattensis var. klamathensis, P. plattensis var. millefolia | P. cascadensis, P. drummondii var. cascadensis |
Name authority | Rydberg: Bull. Torrey Bot. Club 23: 433, plate 277, figs. 1–5. (1896) | Lehmann: Nov. Stirp. Pug. 2: 9. (1830) |
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