Potentilla millefolia |
Potentilla bruceae |
|
---|---|---|
cut-leaf cinquefoil, feather cinquefoil, feather or many-leaf or Klamath cinquefoil, many leaf cinquefoil |
Bruce's cinquefoil |
|
Habit | Plants ± rosetted; taproots fleshy-thickened. | |
Glands | absent or inconspicuous, uncolored. |
|
Stems | usually prostrate, sometimes ± decumbent, 0.4–2(–3) dm, lengths 1–2 times basal leaves. |
ascending to nearly erect, (1–)2–5(–6) dm. |
Basal leaves | pinnate with distal leaflets ± confluent, 2–15(–20) × 1–3 cm; petiole 0.5–2(–3) cm, straight hairs sparse to abundant, appressed to spreading, 0.5–1.5 mm, stiff, cottony hairs absent, glands absent or sparse; primary lateral leaflets (3–)5–13 per side, on distal 2/3–3/4+ of leaf axis, separate to overlapping, largest ones cuneate to flabellate, 0.5–1.5(–2) × 0.5–2 cm, distal 2/3 to whole margin palmately or unevenly, rarely pinnately, incised 2/3 to completely to midvein, ultimate teeth or segments (1–)2–10, linear to broadly oblanceolate, 2–10 × (0.5–)1–2 mm, apical tufts to 1 mm, surfaces green to grayish green, not glaucous, straight hairs sparse to abundant, appressed to spreading, 0.5–1.5(–2) mm, stiff, cottony hairs absent, glands sparse to common. |
not in ranks, ± subpalmate to subpinnate (proximal leaflets sometimes doubled, distal leaflets confluent and/or decurrent), (2–)5–20(–28) cm; petiole (1–)3–15(–20) cm, long hairs abundant, appressed to ascending-spreading, 1–2.5 mm, weak often flattened and twisted, sometimes grading to crisped or cottony hairs, short hairs absent, glands absent or sparse; leaflets 5–7, on distal 1/10–1/4 of leaf axis, ± overlapping, largest ones usually ± obovate, (1–)1.5–5(–6) × 1–3.5(–4) cm, margins flat, distal 2/3–3/4 evenly to unevenly incised ± 1/2 to midvein (often with additional incisions nearly to midvein), undivided medial blade 4–20 mm wide, teeth (2–)3–6(–8) per side (sometimes secondarily toothed), ± lanceolate, 4–11 mm, surfaces ± similar, abaxial ± lighter and hairier, gray-green to grayish, rarely green, not glaucous, long hairs ± abundant, short-crisped hairs absent or sparse to common, cottony hairs usually absent, glands absent or sparse. |
Cauline leaves | (0–)1–2. |
1–2. |
Inflorescences | 3–6(–10)-flowered, loosely cymose, sometimes racemiform. |
5–50-flowered. |
Pedicels | (0.5–)1–2(–4.5) cm, ± recurved in fruit. |
0.5–3(–4) cm. |
Flowers | epicalyx bractlets ± elliptic, 2–4(–6) × 1–2(–2.5) mm; hypanthium 3–6 mm diam.; sepals 4–6(–8) mm, apex acute; petals 4–8(–10) × 3–7(–9) mm; filaments 2–3.5 mm, anthers 0.7–1 mm; carpels 10–30, styles (1.5–)2–3 mm. |
epicalyx bractlets broadly lanceolate to elliptic, 3–6(–7.5) × 1.5–2.5(–3) mm, hairs ± common to abundant, loosely appressed to ascending, glands absent or inconspicuous; hypanthium 3.5–5.5 mm diam.; sepals 5–10 mm, apex acute to acuminate; petals 5–10 × 5–10 mm; filaments 1–2.5 mm, anthers 0.7–1.2 mm; carpels 15–30, styles filiform-tapered, papillate-swollen proximally, 2–3 mm. |
Achenes | 1.5–2 mm, smooth, often ± carunculate. |
1.4–1.6 mm. |
2n | = 63–73, 79, 87, 98, 129. |
|
Potentilla millefolia |
Potentilla bruceae |
|
Phenology | Flowering spring–summer. | Flowering summer. |
Habitat | Vernally to permanently wet meadows, moist openings in conifer forests and sagebrush, alkaline flats | Dry montane meadows and adjacent slopes, in conifer woodlands |
Elevation | 700–2200 m (2300–7200 ft) | 1200–3700 m (3900–12100 ft) |
Distribution |
CA; NV; OR
|
CA; NV; OR
|
Discussion | Potentilla millefolia occurs from central Oregon to the east side of the Sierra Nevada of California, with a disjunct occurrence on the alkaline flats of Reese River Valley, Nevada. Significant variation occurs in vestiture type, leaflet dissection, and flower size, but with minimal geographic correlation. The most distinctive variant, represented by the type of P. klamathensis, has relatively long, slender, spreading, pustule-based hairs, often intermixed with shorter hairs. This vestiture type does not appear to be correlated with any other characters or geographic distribution and may vary within a population. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
As redefined by B. Ertter and D. Mansfield (2007), Potentilla bruceae is characterized by irregularly subpalmate, weakly villose leaves. Such plants form relatively consistent and extensive populations in dry meadows centered near Lake Tahoe in California and Nevada, extending to the central Sierra Nevada and to the Warner Mountains in Oregon. This interpretation differs from that previously suggested by Ertter (1992), in which P. bruceae was interpreted as the catchall category for hybrids between P. breweri (sect. Multijugae) and P. drummondii. All three taxa were accordingly treated as varieties of P. drummondii. D. D. Keck (in J. Clausen et al. 1940) treated P. bruceae as a subspecies of P. drummondii; B. C. Johnston (1980) concluded that its placement as a variety of P. breweri was more justified. The three entities unquestionably intergrade (with the type of P. anomalofolia M. Peck possibly one result), but not beyond what is the norm for facultatively apomictic species of Potentilla. Chromosome numbers reported by J. Clausen et al. (1940) require verification as this species; the irregular behavior observed in meiotic material suggests that at least some of the individuals were hybrids. See discussion of 29. Potentilla drummondii regarding leaflet dissection and the inclusion of P. bruceae in sect. Graciles. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 9, p. 174. | FNA vol. 9, p. 159. |
Parent taxa | Rosaceae > subfam. Rosoideae > tribe Potentilleae > Potentilla > sect. Multijugae | Rosaceae > subfam. Rosoideae > tribe Potentilleae > Potentilla > sect. Graciles |
Sibling taxa | ||
Synonyms | P. klamathensis, P. millefolia var. klamathensis, P. plattensis var. klamathensis, P. plattensis var. millefolia | P. drummondii subsp. bruceae, P. drummondii var. bruceae |
Name authority | Rydberg: Bull. Torrey Bot. Club 23: 433, plate 277, figs. 1–5. (1896) | Rydberg: in N. L. Britton et al., N. Amer. Fl. 22: 342. (1908) |
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