Potentilla millefolia |
Potentilla basaltica |
|
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cut-leaf cinquefoil, feather cinquefoil, feather or many-leaf or Klamath cinquefoil, many leaf cinquefoil |
Black Rock potentilla, Soldier Meadow or Black Rock cinquefoil, soldier meadows cinquefoil |
|
Habit | Plants ± rosetted; taproots fleshy-thickened. | Plants ± rosetted; taproots fleshy-thickened. |
Stems | usually prostrate, sometimes ± decumbent, 0.4–2(–3) dm, lengths 1–2 times basal leaves. |
± prostrate, sometimes ascending in supporting vegetation, 1.5–5 dm, lengths 2–3 times basal leaves. |
Basal leaves | pinnate with distal leaflets ± confluent, 2–15(–20) × 1–3 cm; petiole 0.5–2(–3) cm, straight hairs sparse to abundant, appressed to spreading, 0.5–1.5 mm, stiff, cottony hairs absent, glands absent or sparse; primary lateral leaflets (3–)5–13 per side, on distal 2/3–3/4+ of leaf axis, separate to overlapping, largest ones cuneate to flabellate, 0.5–1.5(–2) × 0.5–2 cm, distal 2/3 to whole margin palmately or unevenly, rarely pinnately, incised 2/3 to completely to midvein, ultimate teeth or segments (1–)2–10, linear to broadly oblanceolate, 2–10 × (0.5–)1–2 mm, apical tufts to 1 mm, surfaces green to grayish green, not glaucous, straight hairs sparse to abundant, appressed to spreading, 0.5–1.5(–2) mm, stiff, cottony hairs absent, glands sparse to common. |
pinnate with distal leaflets ± confluent, 5–12(–18) × 1–1.5(–2) cm; petiole 0.5–1(–1.5) cm, straight hairs absent, cottony hairs absent, glands absent or sparse; primary lateral leaflets (5–)10–15(–20) per side, on nearly whole leaf axis, ± overlapping, largest ones (or leaflet lobes) elliptic, 0.4–1(–2) × 0.2–0.5(–1) cm, simple and entire or 2(–3)-fid to base (sometimes shallowly toothed as well), segments 1–2(–3), narrowly to broadly elliptic, 4–10(–20) × (1–)2–3 mm, apical tufts absent or less than 0.5 mm, surfaces green, glaucous, straight hairs absent or sparse to common, tightly appressed, 0.5 mm or less, stiff, cottony hairs absent, glands absent or sparse. |
Cauline leaves | (0–)1–2. |
2–4. |
Inflorescences | 3–6(–10)-flowered, loosely cymose, sometimes racemiform. |
(5–)10–20-flowered, loosely cymose. |
Pedicels | (0.5–)1–2(–4.5) cm, ± recurved in fruit. |
0.8–3(–4) cm, straight to ± recurved in fruit. |
Flowers | epicalyx bractlets ± elliptic, 2–4(–6) × 1–2(–2.5) mm; hypanthium 3–6 mm diam.; sepals 4–6(–8) mm, apex acute; petals 4–8(–10) × 3–7(–9) mm; filaments 2–3.5 mm, anthers 0.7–1 mm; carpels 10–30, styles (1.5–)2–3 mm. |
epicalyx bractlets lanceolate-elliptic to ovate-elliptic, 1.5–2.5(–3) × 1 mm; hypanthium 2–5 mm diam.; sepals (3–)4–5(–6) mm, apex acute to acuminate; petals (3.5–)4.5–6.5 × 2.5–4 mm; filaments 1.5–3 mm, anthers 0.6–0.8 mm; carpels 3–10, styles 2–2.5 mm. |
Achenes | 1.5–2 mm, smooth, often ± carunculate. |
1.8–2.2 mm, smooth to ± rugose, not carunculate. |
Potentilla millefolia |
Potentilla basaltica |
|
Phenology | Flowering spring–summer. | Flowering late spring–summer. |
Habitat | Vernally to permanently wet meadows, moist openings in conifer forests and sagebrush, alkaline flats | Moist, subalkaline meadows in shrub steppe |
Elevation | 700–2200 m (2300–7200 ft) | 1300–1600 m (4300–5200 ft) |
Distribution |
CA; NV; OR
|
CA; NV |
Discussion | Potentilla millefolia occurs from central Oregon to the east side of the Sierra Nevada of California, with a disjunct occurrence on the alkaline flats of Reese River Valley, Nevada. Significant variation occurs in vestiture type, leaflet dissection, and flower size, but with minimal geographic correlation. The most distinctive variant, represented by the type of P. klamathensis, has relatively long, slender, spreading, pustule-based hairs, often intermixed with shorter hairs. This vestiture type does not appear to be correlated with any other characters or geographic distribution and may vary within a population. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. Potentilla basaltica is very distinctive in its glaucous, highly divided cylindric leaves with leaflets arranged in four ranks. The leaves strongly resemble those of some species of Ivesia, especially I. kingii S. Watson, but molecular analysis (B. Ertter et al. 1998) confirms a relationship with other members of sect. Multijugae. Potentilla basaltica is known from only two localities, one in Lassen County, California, and the other in Humboldt County, Nevada. Both populations are highly localized, and P. basaltica is a candidate for federal listing as well as of conservation concern in both states. The epithet basaltica was chosen as a reference to the Black Rock Desert, not habitat. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 9, p. 174. | FNA vol. 9, p. 176. |
Parent taxa | Rosaceae > subfam. Rosoideae > tribe Potentilleae > Potentilla > sect. Multijugae | Rosaceae > subfam. Rosoideae > tribe Potentilleae > Potentilla > sect. Multijugae |
Sibling taxa | ||
Synonyms | P. klamathensis, P. millefolia var. klamathensis, P. plattensis var. klamathensis, P. plattensis var. millefolia | |
Name authority | Rydberg: Bull. Torrey Bot. Club 23: 433, plate 277, figs. 1–5. (1896) | Tiehm & Ertter: Brittonia 36: 228, fig. 1. (1984) |
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