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cut-leaf cinquefoil, feather cinquefoil, feather or many-leaf or Klamath cinquefoil, many leaf cinquefoil

common silverweed, Pacific cinquefoil, silver weed cinquefoil, silverweed

Habit Plants ± rosetted; taproots fleshy-thickened.
Stems

usually prostrate, sometimes ± decumbent, 0.4–2(–3) dm, lengths 1–2 times basal leaves.

Basal leaves

pinnate with distal leaflets ± confluent, 2–15(–20) × 1–3 cm;

petiole 0.5–2(–3) cm, straight hairs sparse to abundant, appressed to spreading, 0.5–1.5 mm, stiff, cottony hairs absent, glands absent or sparse;

primary lateral leaflets (3–)5–13 per side, on distal 2/3–3/4+ of leaf axis, separate to overlapping, largest ones cuneate to flabellate, 0.5–1.5(–2) × 0.5–2 cm, distal 2/3 to whole margin palmately or unevenly, rarely pinnately, incised 2/3 to completely to midvein, ultimate teeth or segments (1–)2–10, linear to broadly oblanceolate, 2–10 × (0.5–)1–2 mm, apical tufts to 1 mm, surfaces green to grayish green, not glaucous, straight hairs sparse to abundant, appressed to spreading, 0.5–1.5(–2) mm, stiff, cottony hairs absent, glands sparse to common.

petiole (0.5–)1–15(–25) cm, long hairs absent or sparse to dense, 1.5–3.5 mm;

larger leaflets (0.4–)0.5–5(–7) × (0.2–)0.3–2(–3) cm, surfaces: abaxial with long hairs absent or sparse to abundant, not restricted to veins, 0.5–2(–2.5) mm, cottony-crisped hairs usually dense or usually absent (in subsp. groenlandica), adaxial with long hairs absent or sparse to abundant, cottony-crisped hairs usually absent, sometimes sparse to common (especially subsp. yukonensis).

Cauline leaves

(0–)1–2.

Inflorescences

3–6(–10)-flowered, loosely cymose, sometimes racemiform.

Pedicels

(0.5–)1–2(–4.5) cm, ± recurved in fruit.

Flowers

epicalyx bractlets ± elliptic, 2–4(–6) × 1–2(–2.5) mm;

hypanthium 3–6 mm diam.;

sepals 4–6(–8) mm, apex acute;

petals 4–8(–10) × 3–7(–9) mm;

filaments 2–3.5 mm, anthers 0.7–1 mm;

carpels 10–30, styles (1.5–)2–3 mm.

epicalyx bractlets narrowly to broadly ovate-triangular or linear to elliptic, (2–)2.5–7(–8) × (0.3–)0.5–3(–3.5) mm, often 2-fid or dentate;

sepals (3–)3.5–7(–9) mm, apex subacute to acuminate;

petals (4–)5–15(–20) × (2.5–)3–10(–12) mm;

filaments (1–)2.5–3.5(–4.5) mm, anthers 0.7–1.3 mm;

carpels (10–)20–200(–250).

Achenes

1.5–2 mm, smooth, often ± carunculate.

2 mm.

Potentilla millefolia

Potentilla anserina

Phenology Flowering spring–summer.
Habitat Vernally to permanently wet meadows, moist openings in conifer forests and sagebrush, alkaline flats
Elevation 700–2200 m (2300–7200 ft)
Distribution
from FNA
CA; NV; OR
[WildflowerSearch map]
[BONAP county map]
from FNA
AK; AZ; CA; CO; CT; IA; ID; IL; IN; MA; ME; MI; MN; MT; ND; NE; NH; NM; NV; NY; OH; OR; RI; SD; UT; VT; WA; WI; WY; AB; BC; MB; NB; NL; NS; NT; NU; ON; PE; QC; SK; YT; Mexico; South America (Argentina, Chile); Eurasia; Pacific Islands (New Guinea, New Zealand); s Australia
[WildflowerSearch map]
[BONAP county map]
Discussion

Potentilla millefolia occurs from central Oregon to the east side of the Sierra Nevada of California, with a disjunct occurrence on the alkaline flats of Reese River Valley, Nevada. Significant variation occurs in vestiture type, leaflet dissection, and flower size, but with minimal geographic correlation. The most distinctive variant, represented by the type of P. klamathensis, has relatively long, slender, spreading, pustule-based hairs, often intermixed with shorter hairs. This vestiture type does not appear to be correlated with any other characters or geographic distribution and may vary within a population.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Subspecies 4 or 5 (4 in the flora).

Potentilla anserina is polymorphic in most features, especially in hairiness, but also in size and in degree of dissection of leaflets, epicalyx bractlets, and sepals. While most of the variation described by A. G. Blytt (1906) is taxonomically insignificant, A. Rousi (1965) found support for three northern races: subspp. anserina, egedei, and pacifica. He also suggested that P. yukonensis Hultén might qualify as a separate subspecies. This treatment follows J. Soják (1994) in accepting four northern subspecies of P. anserina, all present in North America.

A. Rousi (1965) demonstrated partial interfertility between the races of Potentilla anserina, which form intermediates in all zones of contact. The distinctness of the four races, in spite of intermediates, is upheld by being partly allopatric and occupying different habitats where they are sympatric. As partly interfertile parapatric entities, they are best treated as subspecies.

The vast majority of chromosome counts are tetraploid. Tetraploid plants (2n = 28) are fully fertile; hexaploids (2n = 42) are largely pollen and seed sterile; pentaploids (2n = 35) are probably occasional hybrids (S. Erlandsson 1942, 1942b; A. Rousi 1965). A. Kurtto et al. (in J. Jalas et al. 1972+, vol. 13) considered numbers above tetraploid level as cases of occasional autopolyploidy.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Epicalyx bractlets equal to or longer than sepals, often 2-fid or dentate; leaflet surfaces: abaxial with cottony-crisped hairs dense, long hairs common to abundant, on and between veins, adaxial glabrous or sparsely to densely hairy; achenes with dorsal groove; inland and seashore plants
→ 2
1. Epicalyx bractlets shorter than sepals, usually entire, rarely 2-fid or dentate; leaflet surfaces: abaxial with cottony-crisped hairs absent or sparse to dense, long hairs absent or sparse, on veins, adaxial usually glabrous, rarely sparsely to densely hairy; achenes without dorsal groove; seashore or near-coastal plants
→ 3
2. Hypanthium patelliform (wider than deep) in fruit; epicalyx bractlets narrowly to broadly ovate-triangular, equal to sepals; most of North America.
subsp. anserina
2. Hypanthium turbinate (± as deep as wide) in fruit; epicalyx bractlets usually linear to elliptic, longer than sepals, rarely subequal; interior w Canada and Alaska.
subsp. yukonensis
3. Carpels (20–)50–200(–250); leaves (3–)10–50(–75) cm; leaflets (4–)5–10(–15) per side, abaxial surfaces densely hairy, teeth (4–)6–12(–16) per side, teeth apices acute to acuminate, rarely subacute; flowers 1–2.5(–3.5) cm diam.
subsp. pacifica
3. Carpels 25–60; leaves (1–)2–10 cm, rarely longer; leaflets 2–4(–5) per side, abaxial surfaces glabrous, sometimes sparsely to densely hairy, teeth 2–6(–10) per side, teeth apices rounded to subacute; flowers 0.8–1.5 cm diam.
subsp. groenlandica
Source FNA vol. 9, p. 174. FNA vol. 9, p. 127.
Parent taxa Rosaceae > subfam. Rosoideae > tribe Potentilleae > Potentilla > sect. Multijugae Rosaceae > subfam. Rosoideae > tribe Potentilleae > Potentilla > sect. Pentaphylloides
Sibling taxa
P. albiflora, P. ambigens, P. anachoretica, P. angelliae, P. anglica, P. anserina, P. arenosa, P. argentea, P. arizonica, P. basaltica, P. bicrenata, P. biennis, P. biflora, P. bimundorum, P. bipinnatifida, P. brevifolia, P. breweri, P. bruceae, P. brunnescens, P. canadensis, P. concinna, P. cottamii, P. crantzii, P. crebridens, P. crinita, P. cristae, P. demotica, P. drummondii, P. effusa, P. elegans, P. erecta, P. flabellifolia, P. fragiformis, P. furcata, P. glaucophylla, P. gracilis, P. grayi, P. hickmanii, P. hippiana, P. holmgrenii, P. hookeriana, P. hyparctica, P. inclinata, P. intermedia, P. jepsonii, P. johnstonii, P. lasiodonta, P. litoralis, P. macounii, P. modesta, P. morefieldii, P. multijuga, P. multisecta, P. nana, P. newberryi, P. nivea, P. norvegica, P. ovina, P. paucijuga, P. pedersenii, P. pensylvanica, P. plattensis, P. pseudosericea, P. pulchella, P. pulcherrima, P. recta, P. reptans, P. rhyolitica, P. rimicola, P. rivalis, P. robbinsiana, P. rubella, P. rubricaulis, P. sanguinea, P. saximontana, P. sierrae-blancae, P. simplex, P. sterilis, P. stipularis, P. subgorodkovii, P. subjuga, P. subvahliana, P. subviscosa, P. supina, P. thurberi, P. thuringiaca, P. tikhomirovii, P. townsendii, P. uliginosa, P. uschakovii, P. vahliana, P. verna, P. versicolor, P. villosa, P. villosula, P. vulcanicola, P. wheeleri
P. albiflora, P. ambigens, P. anachoretica, P. angelliae, P. anglica, P. arenosa, P. argentea, P. arizonica, P. basaltica, P. bicrenata, P. biennis, P. biflora, P. bimundorum, P. bipinnatifida, P. brevifolia, P. breweri, P. bruceae, P. brunnescens, P. canadensis, P. concinna, P. cottamii, P. crantzii, P. crebridens, P. crinita, P. cristae, P. demotica, P. drummondii, P. effusa, P. elegans, P. erecta, P. flabellifolia, P. fragiformis, P. furcata, P. glaucophylla, P. gracilis, P. grayi, P. hickmanii, P. hippiana, P. holmgrenii, P. hookeriana, P. hyparctica, P. inclinata, P. intermedia, P. jepsonii, P. johnstonii, P. lasiodonta, P. litoralis, P. macounii, P. millefolia, P. modesta, P. morefieldii, P. multijuga, P. multisecta, P. nana, P. newberryi, P. nivea, P. norvegica, P. ovina, P. paucijuga, P. pedersenii, P. pensylvanica, P. plattensis, P. pseudosericea, P. pulchella, P. pulcherrima, P. recta, P. reptans, P. rhyolitica, P. rimicola, P. rivalis, P. robbinsiana, P. rubella, P. rubricaulis, P. sanguinea, P. saximontana, P. sierrae-blancae, P. simplex, P. sterilis, P. stipularis, P. subgorodkovii, P. subjuga, P. subvahliana, P. subviscosa, P. supina, P. thurberi, P. thuringiaca, P. tikhomirovii, P. townsendii, P. uliginosa, P. uschakovii, P. vahliana, P. verna, P. versicolor, P. villosa, P. villosula, P. vulcanicola, P. wheeleri
Subordinate taxa
P. anserina subsp. anserina, P. anserina subsp. groenlandica, P. anserina subsp. pacifica, P. anserina subsp. yukonensis
Synonyms P. klamathensis, P. millefolia var. klamathensis, P. plattensis var. klamathensis, P. plattensis var. millefolia Argentina anserina
Name authority Rydberg: Bull. Torrey Bot. Club 23: 433, plate 277, figs. 1–5. (1896) Linnaeus: Sp. Pl. 1: 495. (1753)
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