Potentilla hippiana
Potentilla vulcanicola
Hipp's cinquefoil, horse cinquefoil, potentille de Hipp, woolly cinquefoil
stout, sometimes ± columnar, not sheathed with marcescent whole leaves.
(0.3–)2–4(–5) dm, lengths (1–)2–4 times basal leaves.
ascending to erect, (0.2–)0.4–1.6(–2) dm, lengths 2–3 times basal leaves.
pinnate to subpinnate, (3–)5–15(–25) cm;
petiole 1–10(–15) cm, long hairs abundant to dense, tightly appressed, 1–2.5 mm, stiff, short hairs absent or obscured, crisped-cottony hairs absent or sparse to common, glands absent or obscured;
leaflets not conduplicate, lateral ones evenly (to unevenly in argyrea phase) paired, (2–)3–6(–7) per side on distal (1/6–)1/5–1/2 of leaf axis, distal pairs ± decurrent, often confluent with terminal leaflet, larger leaflets oblanceolate or narrowly obovate to oblong, 1–5(–6) × 0.3–1.5 cm, distal (2/3–)3/4 to whole margin incised 1/2 or less (rarely more) to midvein, teeth (5–)7–12(–18) per side, 1–4 mm, surfaces ± to strongly dissimilar, abaxial white, long hairs abundant (mostly on, but not limited to, veins), 1–2 mm, stiff, short hairs absent or obscured, crisped-cottony hairs abundant to dense, glands absent or obscured, adaxial green to grayish, long hairs sparse to common, rarely absent, short or crisped to, sometimes, cottony hairs absent or sparse to common, glands sparse.
1–6(–11) cm;
petiole 0.5–4(–9) cm, long hairs ± abundant, ascending to spreading, sometimes loosely appressed, 1–2.5 mm, soft to weak, smooth, crisped(/cottony) hairs absent or sparse, sometimes common, glands absent or sparse;
leaflets overlapping, central broadly obovate, 0.8–2(–2.5) × 0.8–1.5(–2.5) cm, sessile or subsessile, base broadly cuneate, margins strongly revolute, distal 1/2–2/3 incised ± 1/2 to midvein, teeth 2–3(–4) per side, distant, surfaces strongly dissimilar, abaxial grayish white, long hairs 1–2 mm, cottony-crisped hairs ± dense, adaxial usually dark green, sometimes grayish green, long hairs sparse to abundant, other hairs usually absent.
1–2(–3).
(0–)1–2(–3).
10–30-flowered.
1–2(–5)-flowered.
0.3–3(–5) cm.
1–2 cm in flower, to 5 cm in fruit.
epicalyx bractlets narrowly elliptic to lanceolate, rarely linear, 2–5 × 0.5–1.5 mm, (1/2–)2/3 as long as sepals, abaxial vestiture similar to or ± sparser than sepals, usually not glabrescent, straight hairs ± abundant, crisped to sometimes ± cottony hairs absent or sparse to abundant;
hypanthium 3–7 mm diam.;
sepals 4–5.5(–6.5) mm, apex acute to acuminate;
petals 4–8 × 4–7 mm;
filaments 0.5–2.5 mm, anthers 0.6–1.1 mm;
carpels (5–)10–30, styles 1.7–2.5 mm.
epicalyx bractlets ± oblong to ovate, 4–8(–10) × 1.8–3(–5) mm, 2/3 to as wide as sepals, margins revolute, sometimes flat, red glands absent;
hypanthium 3.5–4.5 mm diam.;
sepals 5–8 mm, apex ± acute;
petals 7–9 × 7–10 mm, longer than sepals;
filaments 0.8–1 mm, anthers 0.6–0.7 mm;
carpels 20–30, apical hairs sparse to abundant (straight), styles conic-columnar, ± papillate-swollen in less than proximal 1/5, or sometimes to 1/5, 0.8–1.1 mm.
1.4–1.8 mm, smooth to faintly rugose.
0.9–1.2 mm.
= 42, 70, 77, 84, 98.
= 28 (Russian Far East).
Potentilla hippiana
Potentilla vulcanicola
Potentilla hippiana is most common and distinctive in the Colorado Plateau and southern Rocky Mountains, with outliers in the mountains of Arizona, Nevada, and New Mexico. Occurrences are more scattered in the northern Rocky Mountains and Great Plains, where intergradation with P. effusa is common. Among these intermediates are the argyrea phase and the single report from Idaho (B. C. Johnston 1980). Potentilla hippiana is probably adventive in Nova Scotia, eastern Ontario, Quebec, and Michigan, and possibly the Northwest Territories. The reported occurrence in central Alaska (E. Hultén 1968) is based on a specimen of P. pensylvanica (CAN).
Significant variation exists throughout the range of Potentilla hippiana, including dwarfism, leaf division, leaflet dissection, adaxial leaflet vestiture, and pedicel length. Some of this variation might merit taxonomic recognition upon further analysis, especially if correlated with ploidy level (6x through 12x). At present, the differences used to distinguish var. argyrea are not found to be sufficiently correlated with distribution to merit formal taxonomic recognition.
Complicating the infraspecific variation is a tendency for Potentilla hippiana to intergrade with other species, in particular P. effusa. Beyond sect. Leucophyllae, the frequency of hybrids with P. pulcherrima blurs the distinction between the two species; P. gracilis var. hippianoides S. L. Welsh & N. D. Atwood is probably one of the resultant intermediates.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Potentilla vulcanicola is close morphologically to P. uniflora in the narrow sense and largely replaces it in southern and eastern Chukotka, Alaska, and northwestern Canada, where it reaches east of Mackenzie River on the coastal mainland and the westernmost islands (Banks and Victoria islands). Potentilla subgorodkovii and P. vulcanicola account for a major part of North American records of P. uniflora.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
BC
CalFlora
MI Flora
MN Wildflowers
PNW Herbaria
WildflowerSearch
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