Potentilla hippiana |
Potentilla uschakovii |
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Hipp's cinquefoil, horse cinquefoil, potentille de Hipp, woolly cinquefoil |
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Caudex branches | often sheathed with marcescent whole leaves. |
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Stems | (0.3–)2–4(–5) dm, lengths (1–)2–4 times basal leaves. |
ascending to nearly erect, 0.3–1.5 dm. |
Basal leaves | pinnate to subpinnate, (3–)5–15(–25) cm; petiole 1–10(–15) cm, long hairs abundant to dense, tightly appressed, 1–2.5 mm, stiff, short hairs absent or obscured, crisped-cottony hairs absent or sparse to common, glands absent or obscured; leaflets not conduplicate, lateral ones evenly (to unevenly in argyrea phase) paired, (2–)3–6(–7) per side on distal (1/6–)1/5–1/2 of leaf axis, distal pairs ± decurrent, often confluent with terminal leaflet, larger leaflets oblanceolate or narrowly obovate to oblong, 1–5(–6) × 0.3–1.5 cm, distal (2/3–)3/4 to whole margin incised 1/2 or less (rarely more) to midvein, teeth (5–)7–12(–18) per side, 1–4 mm, surfaces ± to strongly dissimilar, abaxial white, long hairs abundant (mostly on, but not limited to, veins), 1–2 mm, stiff, short hairs absent or obscured, crisped-cottony hairs abundant to dense, glands absent or obscured, adaxial green to grayish, long hairs sparse to common, rarely absent, short or crisped to, sometimes, cottony hairs absent or sparse to common, glands sparse. |
often both ternate and palmate or subpalmate on same plant, 1–5 cm; petiole 0.5–3 cm, long hairs sparse to dense, ± appressed to ascending, (1–)1.5–2.5 mm, soft to ± weak, smooth, short hairs absent or sparse, crisped(/cottony) hairs sparse to common, glands absent or sparse; leaflets 3–5, proximalmost separated by 0–2 mm, central broadly elliptic to obovate, 0.5–2 × 0.3–1.4 cm, petiolules ± 1 mm, distal 2/3 to nearly whole margin incised 2/3–3/5+ to midvein, teeth (2–)3–4 per side, 3–5 mm, apical tufts 1–1.5 mm, abaxial surfaces grayish white to white, long hairs common to abundant, cottony(/crisped) hairs dense, short hairs and glands absent or obscured, adaxial grayish green to gray, long hairs abundant, 1–2 mm, soft (grading to cottony), short/crisped/cottony hairs common to abundant, glands sparse to common. |
Cauline leaves | 1–2(–3). |
0–1. |
Inflorescences | 10–30-flowered. |
1–3(–4)-flowered, open, branch angle 20–40°. |
Pedicels | 0.3–3(–5) cm. |
1–1.5 cm, proximal to 3 cm. |
Flowers | epicalyx bractlets narrowly elliptic to lanceolate, rarely linear, 2–5 × 0.5–1.5 mm, (1/2–)2/3 as long as sepals, abaxial vestiture similar to or ± sparser than sepals, usually not glabrescent, straight hairs ± abundant, crisped to sometimes ± cottony hairs absent or sparse to abundant; hypanthium 3–7 mm diam.; sepals 4–5.5(–6.5) mm, apex acute to acuminate; petals 4–8 × 4–7 mm; filaments 0.5–2.5 mm, anthers 0.6–1.1 mm; carpels (5–)10–30, styles 1.7–2.5 mm. |
epicalyx bractlets lanceolate to ovate, 3–4 × 1–2 mm; hypanthium 2.5–3.5 mm diam.; sepals 3.5–5 mm, apex obtuse to subacute, glands sparse to common, often ± obscured; petals pale yellow, often overlapping, 5–7 × 5–8(–9) mm, distinctly longer than sepals; filaments 0.5–1.5 mm, anthers ± 0.5 mm; carpels 30–60, styles 0.8–0.9 mm. |
Achenes | 1.4–1.8 mm, smooth to faintly rugose. |
1–1.3 mm. |
2n | = 42, 70, 77, 84, 98. |
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Potentilla hippiana |
Potentilla uschakovii |
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Phenology | Flowering summer. | Flowering summer. |
Habitat | Dry grasslands and meadows, in aspen and conifer woodlands or alpine tundra, disturbed sites | Dry tundra meadows, gravel and loam ridges, loam flats, sandy bluffs |
Elevation | 500–3400 m (1600–11200 ft) | 0–200 m (0–700 ft) |
Distribution |
AZ; CO; ID; MI; MN; MT; ND; NE; NM; NV; SD; UT; WY; AB; BC; MB; NS; NT; ON; QC; SK
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NT; NU; Greenland; e Asia (Russian Far East) |
Discussion | Potentilla hippiana is most common and distinctive in the Colorado Plateau and southern Rocky Mountains, with outliers in the mountains of Arizona, Nevada, and New Mexico. Occurrences are more scattered in the northern Rocky Mountains and Great Plains, where intergradation with P. effusa is common. Among these intermediates are the argyrea phase and the single report from Idaho (B. C. Johnston 1980). Potentilla hippiana is probably adventive in Nova Scotia, eastern Ontario, Quebec, and Michigan, and possibly the Northwest Territories. The reported occurrence in central Alaska (E. Hultén 1968) is based on a specimen of P. pensylvanica (CAN). Significant variation exists throughout the range of Potentilla hippiana, including dwarfism, leaf division, leaflet dissection, adaxial leaflet vestiture, and pedicel length. Some of this variation might merit taxonomic recognition upon further analysis, especially if correlated with ploidy level (6x through 12x). At present, the differences used to distinguish var. argyrea are not found to be sufficiently correlated with distribution to merit formal taxonomic recognition. Complicating the infraspecific variation is a tendency for Potentilla hippiana to intergrade with other species, in particular P. effusa. Beyond sect. Leucophyllae, the frequency of hybrids with P. pulcherrima blurs the distinction between the two species; P. gracilis var. hippianoides S. L. Welsh & N. D. Atwood is probably one of the resultant intermediates. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
For discussion of probable parentage and resultant diagnostic characters, see discussion of 89. Potentilla pedersenii. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 9, p. 162. | FNA vol. 9, p. 211. |
Parent taxa | Rosaceae > subfam. Rosoideae > tribe Potentilleae > Potentilla > sect. Leucophyllae | Rosaceae > subfam. Rosoideae > tribe Potentilleae > Potentilla > sect. Rubricaules |
Sibling taxa | ||
Synonyms | P. leucophylla, P. argyrea, P. effusa var. argyrea, P. hippiana var. argyrea, P. hippiana var. diffusa, P. propinqua | |
Name authority | Lehmann: Nov. Stirp. Pug. 2: 7. (1830) — not Pallas 1773 | Jurtzev: Bot. Zhurn. (Moscow & Leningrad) 73: 1613. (1988) |
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