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Hipp's cinquefoil, horse cinquefoil, potentille de Hipp, woolly cinquefoil

creeping cinquefoil, potentille rampante

Stems

(0.3–)2–4(–5) dm, lengths (1–)2–4 times basal leaves.

soon becoming prostrate, flagelliform, not branched, rooting at some nodes, 1.5–10+ dm.

Basal leaves

pinnate to subpinnate, (3–)5–15(–25) cm;

petiole 1–10(–15) cm, long hairs abundant to dense, tightly appressed, 1–2.5 mm, stiff, short hairs absent or obscured, crisped-cottony hairs absent or sparse to common, glands absent or obscured;

leaflets not conduplicate, lateral ones evenly (to unevenly in argyrea phase) paired, (2–)3–6(–7) per side on distal (1/6–)1/5–1/2 of leaf axis, distal pairs ± decurrent, often confluent with terminal leaflet, larger leaflets oblanceolate or narrowly obovate to oblong, 1–5(–6) × 0.3–1.5 cm, distal (2/3–)3/4 to whole margin incised 1/2 or less (rarely more) to midvein, teeth (5–)7–12(–18) per side, 1–4 mm, surfaces ± to strongly dissimilar, abaxial white, long hairs abundant (mostly on, but not limited to, veins), 1–2 mm, stiff, short hairs absent or obscured, crisped-cottony hairs abundant to dense, glands absent or obscured, adaxial green to grayish, long hairs sparse to common, rarely absent, short or crisped to, sometimes, cottony hairs absent or sparse to common, glands sparse.

persistent, usually palmate, 3–12(–30) cm;

petiole 2–10(–25) cm, long hairs sparse to abundant, tightly to loosely appressed, 0.5–1.5 mm, usually stiff, sometimes weak, glands absent;

leaflets (3–)5(–7), central oblanceolate to obovate, (0.5–)2–4(–7) × 0.3–1.5(–2.5) cm, distal ± 3/4 of margin incised 1/4–1/3 to midvein, teeth (3–)4–12 per side, surfaces similar, green, sparsely to moderately hairy.

Cauline leaves

1–2(–3).

2–3(–4) proximal to 1st flowering node, usually well expanded at anthesis, usually palmate, 2–8(–20) cm;

petiole 0.5–6(–15) cm;

leaflets (3–)5, ± resembling those of basal leaves, apex rounded to obtuse.

Inflorescences

10–30-flowered.

solitary flowers at stolon nodes.

Pedicels

0.3–3(–5) cm.

(2.5–)4–12(–15) cm.

Flowers

epicalyx bractlets narrowly elliptic to lanceolate, rarely linear, 2–5 × 0.5–1.5 mm, (1/2–)2/3 as long as sepals, abaxial vestiture similar to or ± sparser than sepals, usually not glabrescent, straight hairs ± abundant, crisped to sometimes ± cottony hairs absent or sparse to abundant;

hypanthium 3–7 mm diam.;

sepals 4–5.5(–6.5) mm, apex acute to acuminate;

petals 4–8 × 4–7 mm;

filaments 0.5–2.5 mm, anthers 0.6–1.1 mm;

carpels (5–)10–30, styles 1.7–2.5 mm.

5(–10)-merous;

epicalyx bractlets elliptic or oblong to ovate, 4–10 × 1.5–3.5 mm, often much larger than sepals (especially in fruit);

hypanthium 4–7 mm diam.;

sepals (3–)5–7 mm, apex broadly acute to obtuse;

petals 7–9(–12) × 6–9(–11) mm, apex usually ± retuse;

stamens ca. 20, filaments (0.5–)1–2.5(–2.8) mm, anthers (1–)1.3–2 mm;

carpels 60–120, styles 0.6–1.3 mm.

Achenes

1.4–1.8 mm, smooth to faintly rugose.

1.3–1.6 mm, ± rugose.

Rootstocks

erect, slender to stout, 2–6+ cm.

2n

= 42, 70, 77, 84, 98.

= 28 (Eurasia).

Potentilla hippiana

Potentilla reptans

Phenology Flowering summer. Flowering late Apr–Aug.
Habitat Dry grasslands and meadows, in aspen and conifer woodlands or alpine tundra, disturbed sites Dry to moist lawns, roadsides, waste places, on non-acidic soil
Elevation 500–3400 m (1600–11200 ft) 0–500 m (0–1600 ft)
Distribution
from FNA
AZ; CO; ID; MI; MN; MT; ND; NE; NM; NV; SD; UT; WY; AB; BC; MB; NS; NT; ON; QC; SK
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; CO; DC; FL; GA; IL; KY; LA; MA; MD; MI; MN; NJ; NY; OH; OR; PA; VA; WA; WI; NS; ON; QC; Europe; w Asia; n Africa; Atlantic Islands (Azores, Macaronesia) [Introduced in North America; introduced also in West Indies, Bermuda, South America, Africa (Ethiopia), Pacific Islands (New Zealand), Australia]
[BONAP county map]
Discussion

Potentilla hippiana is most common and distinctive in the Colorado Plateau and southern Rocky Mountains, with outliers in the mountains of Arizona, Nevada, and New Mexico. Occurrences are more scattered in the northern Rocky Mountains and Great Plains, where intergradation with P. effusa is common. Among these intermediates are the argyrea phase and the single report from Idaho (B. C. Johnston 1980). Potentilla hippiana is probably adventive in Nova Scotia, eastern Ontario, Quebec, and Michigan, and possibly the Northwest Territories. The reported occurrence in central Alaska (E. Hultén 1968) is based on a specimen of P. pensylvanica (CAN).

Significant variation exists throughout the range of Potentilla hippiana, including dwarfism, leaf division, leaflet dissection, adaxial leaflet vestiture, and pedicel length. Some of this variation might merit taxonomic recognition upon further analysis, especially if correlated with ploidy level (6x through 12x). At present, the differences used to distinguish var. argyrea are not found to be sufficiently correlated with distribution to merit formal taxonomic recognition.

Complicating the infraspecific variation is a tendency for Potentilla hippiana to intergrade with other species, in particular P. effusa. Beyond sect. Leucophyllae, the frequency of hybrids with P. pulcherrima blurs the distinction between the two species; P. gracilis var. hippianoides S. L. Welsh & N. D. Atwood is probably one of the resultant intermediates.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Potentilla reptans is becoming widely introduced in North America, though not fully naturalized in some of the states indicated here. It may be confused with P. canadensis and P. simplex, but, in addition to the characters highlighted in the key, P. reptans tends to have more crenately toothed leaves than the more sharply toothed leaves of the native species. The plant has a history of medicinal uses similar to that of P. erecta. The cultivated double-flowered form is sometimes found established in weedy places.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 9, p. 162. FNA vol. 9, p. 135.
Parent taxa Rosaceae > subfam. Rosoideae > tribe Potentilleae > Potentilla > sect. Leucophyllae Rosaceae > subfam. Rosoideae > tribe Potentilleae > Potentilla > sect. Potentilla
Sibling taxa
P. albiflora, P. ambigens, P. anachoretica, P. angelliae, P. anglica, P. anserina, P. arenosa, P. argentea, P. arizonica, P. basaltica, P. bicrenata, P. biennis, P. biflora, P. bimundorum, P. bipinnatifida, P. brevifolia, P. breweri, P. bruceae, P. brunnescens, P. canadensis, P. concinna, P. cottamii, P. crantzii, P. crebridens, P. crinita, P. cristae, P. demotica, P. drummondii, P. effusa, P. elegans, P. erecta, P. flabellifolia, P. fragiformis, P. furcata, P. glaucophylla, P. gracilis, P. grayi, P. hickmanii, P. holmgrenii, P. hookeriana, P. hyparctica, P. inclinata, P. intermedia, P. jepsonii, P. johnstonii, P. lasiodonta, P. litoralis, P. macounii, P. millefolia, P. modesta, P. morefieldii, P. multijuga, P. multisecta, P. nana, P. newberryi, P. nivea, P. norvegica, P. ovina, P. paucijuga, P. pedersenii, P. pensylvanica, P. plattensis, P. pseudosericea, P. pulchella, P. pulcherrima, P. recta, P. reptans, P. rhyolitica, P. rimicola, P. rivalis, P. robbinsiana, P. rubella, P. rubricaulis, P. sanguinea, P. saximontana, P. sierrae-blancae, P. simplex, P. sterilis, P. stipularis, P. subgorodkovii, P. subjuga, P. subvahliana, P. subviscosa, P. supina, P. thurberi, P. thuringiaca, P. tikhomirovii, P. townsendii, P. uliginosa, P. uschakovii, P. vahliana, P. verna, P. versicolor, P. villosa, P. villosula, P. vulcanicola, P. wheeleri
P. albiflora, P. ambigens, P. anachoretica, P. angelliae, P. anglica, P. anserina, P. arenosa, P. argentea, P. arizonica, P. basaltica, P. bicrenata, P. biennis, P. biflora, P. bimundorum, P. bipinnatifida, P. brevifolia, P. breweri, P. bruceae, P. brunnescens, P. canadensis, P. concinna, P. cottamii, P. crantzii, P. crebridens, P. crinita, P. cristae, P. demotica, P. drummondii, P. effusa, P. elegans, P. erecta, P. flabellifolia, P. fragiformis, P. furcata, P. glaucophylla, P. gracilis, P. grayi, P. hickmanii, P. hippiana, P. holmgrenii, P. hookeriana, P. hyparctica, P. inclinata, P. intermedia, P. jepsonii, P. johnstonii, P. lasiodonta, P. litoralis, P. macounii, P. millefolia, P. modesta, P. morefieldii, P. multijuga, P. multisecta, P. nana, P. newberryi, P. nivea, P. norvegica, P. ovina, P. paucijuga, P. pedersenii, P. pensylvanica, P. plattensis, P. pseudosericea, P. pulchella, P. pulcherrima, P. recta, P. rhyolitica, P. rimicola, P. rivalis, P. robbinsiana, P. rubella, P. rubricaulis, P. sanguinea, P. saximontana, P. sierrae-blancae, P. simplex, P. sterilis, P. stipularis, P. subgorodkovii, P. subjuga, P. subvahliana, P. subviscosa, P. supina, P. thurberi, P. thuringiaca, P. tikhomirovii, P. townsendii, P. uliginosa, P. uschakovii, P. vahliana, P. verna, P. versicolor, P. villosa, P. villosula, P. vulcanicola, P. wheeleri
Synonyms P. leucophylla, P. argyrea, P. effusa var. argyrea, P. hippiana var. argyrea, P. hippiana var. diffusa, P. propinqua
Name authority Lehmann: Nov. Stirp. Pug. 2: 7. (1830) — not Pallas 1773 Linnaeus: Sp. Pl. 1: 499. (1753)
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