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Hipp's cinquefoil, horse cinquefoil, potentille de Hipp, woolly cinquefoil

Platte cinquefoil, Platte River cinquefoil

Habit Plants rosetted to tufted; taproots fleshy-thickened.
Stems

(0.3–)2–4(–5) dm, lengths (1–)2–4 times basal leaves.

initially decumbent to sometimes ascending, becoming prostrate or supported by vegetation, (0.3–)0.5–3.5(–4.5) dm, lengths 1.5–2.5(–4) times basal leaves.

Basal leaves

pinnate to subpinnate, (3–)5–15(–25) cm;

petiole 1–10(–15) cm, long hairs abundant to dense, tightly appressed, 1–2.5 mm, stiff, short hairs absent or obscured, crisped-cottony hairs absent or sparse to common, glands absent or obscured;

leaflets not conduplicate, lateral ones evenly (to unevenly in argyrea phase) paired, (2–)3–6(–7) per side on distal (1/6–)1/5–1/2 of leaf axis, distal pairs ± decurrent, often confluent with terminal leaflet, larger leaflets oblanceolate or narrowly obovate to oblong, 1–5(–6) × 0.3–1.5 cm, distal (2/3–)3/4 to whole margin incised 1/2 or less (rarely more) to midvein, teeth (5–)7–12(–18) per side, 1–4 mm, surfaces ± to strongly dissimilar, abaxial white, long hairs abundant (mostly on, but not limited to, veins), 1–2 mm, stiff, short hairs absent or obscured, crisped-cottony hairs abundant to dense, glands absent or obscured, adaxial green to grayish, long hairs sparse to common, rarely absent, short or crisped to, sometimes, cottony hairs absent or sparse to common, glands sparse.

pinnate with distal leaflets ± distinct, 2–15(–20) × 1–3 cm;

petiole 0.5–5 cm, straight hairs common, tightly appressed, 0.5(–1) mm, stiff, cottony hairs absent, glands absent or sparse;

primary lateral leaflets (3–)4–6(–8) per side, on distal (1/4–)1/2–3/4 of leaf axis, overlapping to separate, largest ones obovate, 0.5–1.5(–2) × 0.5–1(–1.3) cm, distal (1/2–)2/3 to whole margin pinnately incised 3/4+ to midvein, ultimate teeth 5–10, linear-oblanceolate, 1.5–8(–10) × 1–2 mm, apical tufts less than 0.5 mm, surfaces green to grayish green, not glaucous, straight hairs sparse to common (sparser adaxially), tightly appressed, 0.5 mm, stiff, cottony hairs absent, glands absent or inconspicuous.

Cauline leaves

1–2(–3).

1–3.

Inflorescences

10–30-flowered.

(1–)3–15(–20)-flowered, loosely cymose, sometimes racemiform.

Pedicels

0.3–3(–5) cm.

1–4(–5) cm, ± recurved in fruit.

Flowers

epicalyx bractlets narrowly elliptic to lanceolate, rarely linear, 2–5 × 0.5–1.5 mm, (1/2–)2/3 as long as sepals, abaxial vestiture similar to or ± sparser than sepals, usually not glabrescent, straight hairs ± abundant, crisped to sometimes ± cottony hairs absent or sparse to abundant;

hypanthium 3–7 mm diam.;

sepals 4–5.5(–6.5) mm, apex acute to acuminate;

petals 4–8 × 4–7 mm;

filaments 0.5–2.5 mm, anthers 0.6–1.1 mm;

carpels (5–)10–30, styles 1.7–2.5 mm.

epicalyx bractlets narrowly elliptic to linear-lanceolate, 2–5(–6) × 0.5–2 mm;

hypanthium 3–4 mm diam.;

sepals 3–6 mm, apex acute;

petals 4–7 × 3–6 mm;

filaments 1–2.5 mm, anthers 0.5–1 mm, usually ± 1/2 as long as filaments;

carpels 10–20, styles 1.5–2.5 mm.

Achenes

1.4–1.8 mm, smooth to faintly rugose.

(1.3–)1.5–1.9 mm, smooth, often ± carunculate.

2n

= 42, 70, 77, 84, 98.

= 70.

Potentilla hippiana

Potentilla plattensis

Phenology Flowering summer. Flowering summer.
Habitat Dry grasslands and meadows, in aspen and conifer woodlands or alpine tundra, disturbed sites Moist meadows, streamsides, reservoir margins
Elevation 500–3400 m (1600–11200 ft) 300–2900 m (1000–9500 ft)
Distribution
from FNA
AZ; CO; ID; MI; MN; MT; ND; NE; NM; NV; SD; UT; WY; AB; BC; MB; NS; NT; ON; QC; SK
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; CO; ID; MT; ND; NM; SD; UT; WY; AB; MB; NT; SK
[WildflowerSearch map]
[BONAP county map]
Discussion

Potentilla hippiana is most common and distinctive in the Colorado Plateau and southern Rocky Mountains, with outliers in the mountains of Arizona, Nevada, and New Mexico. Occurrences are more scattered in the northern Rocky Mountains and Great Plains, where intergradation with P. effusa is common. Among these intermediates are the argyrea phase and the single report from Idaho (B. C. Johnston 1980). Potentilla hippiana is probably adventive in Nova Scotia, eastern Ontario, Quebec, and Michigan, and possibly the Northwest Territories. The reported occurrence in central Alaska (E. Hultén 1968) is based on a specimen of P. pensylvanica (CAN).

Significant variation exists throughout the range of Potentilla hippiana, including dwarfism, leaf division, leaflet dissection, adaxial leaflet vestiture, and pedicel length. Some of this variation might merit taxonomic recognition upon further analysis, especially if correlated with ploidy level (6x through 12x). At present, the differences used to distinguish var. argyrea are not found to be sufficiently correlated with distribution to merit formal taxonomic recognition.

Complicating the infraspecific variation is a tendency for Potentilla hippiana to intergrade with other species, in particular P. effusa. Beyond sect. Leucophyllae, the frequency of hybrids with P. pulcherrima blurs the distinction between the two species; P. gracilis var. hippianoides S. L. Welsh & N. D. Atwood is probably one of the resultant intermediates.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Potentilla plattensis occurs mostly east of the Continental Divide from the Canadian Prairies to the mountains of New Mexico. The species barely enters Idaho at Monida Pass (Clark County). Populations also exist in the White Mountains of east-central Arizona and the mountains of southern Utah. The species is relatively uniform throughout its range but there is often significant seasonal variation, such that plants can be compact and densely strigose in early summer but elongate and subglabrous later in the season.

Where the ranges of Potentilla plattensis and P. ovina overlap, the two are sometimes difficult to distinguish. The habitats are usually distinct, in that P. plattensis generally occurs in moist valley bottoms and P. ovina occurs in rocky uplands. Differences in vestiture are also diagnostic: hairs of P. plattensis are 0.5 mm and tightly appressed; those of P. ovina are longer and looser.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 9, p. 162. FNA vol. 9, p. 173.
Parent taxa Rosaceae > subfam. Rosoideae > tribe Potentilleae > Potentilla > sect. Leucophyllae Rosaceae > subfam. Rosoideae > tribe Potentilleae > Potentilla > sect. Multijugae
Sibling taxa
P. albiflora, P. ambigens, P. anachoretica, P. angelliae, P. anglica, P. anserina, P. arenosa, P. argentea, P. arizonica, P. basaltica, P. bicrenata, P. biennis, P. biflora, P. bimundorum, P. bipinnatifida, P. brevifolia, P. breweri, P. bruceae, P. brunnescens, P. canadensis, P. concinna, P. cottamii, P. crantzii, P. crebridens, P. crinita, P. cristae, P. demotica, P. drummondii, P. effusa, P. elegans, P. erecta, P. flabellifolia, P. fragiformis, P. furcata, P. glaucophylla, P. gracilis, P. grayi, P. hickmanii, P. holmgrenii, P. hookeriana, P. hyparctica, P. inclinata, P. intermedia, P. jepsonii, P. johnstonii, P. lasiodonta, P. litoralis, P. macounii, P. millefolia, P. modesta, P. morefieldii, P. multijuga, P. multisecta, P. nana, P. newberryi, P. nivea, P. norvegica, P. ovina, P. paucijuga, P. pedersenii, P. pensylvanica, P. plattensis, P. pseudosericea, P. pulchella, P. pulcherrima, P. recta, P. reptans, P. rhyolitica, P. rimicola, P. rivalis, P. robbinsiana, P. rubella, P. rubricaulis, P. sanguinea, P. saximontana, P. sierrae-blancae, P. simplex, P. sterilis, P. stipularis, P. subgorodkovii, P. subjuga, P. subvahliana, P. subviscosa, P. supina, P. thurberi, P. thuringiaca, P. tikhomirovii, P. townsendii, P. uliginosa, P. uschakovii, P. vahliana, P. verna, P. versicolor, P. villosa, P. villosula, P. vulcanicola, P. wheeleri
P. albiflora, P. ambigens, P. anachoretica, P. angelliae, P. anglica, P. anserina, P. arenosa, P. argentea, P. arizonica, P. basaltica, P. bicrenata, P. biennis, P. biflora, P. bimundorum, P. bipinnatifida, P. brevifolia, P. breweri, P. bruceae, P. brunnescens, P. canadensis, P. concinna, P. cottamii, P. crantzii, P. crebridens, P. crinita, P. cristae, P. demotica, P. drummondii, P. effusa, P. elegans, P. erecta, P. flabellifolia, P. fragiformis, P. furcata, P. glaucophylla, P. gracilis, P. grayi, P. hickmanii, P. hippiana, P. holmgrenii, P. hookeriana, P. hyparctica, P. inclinata, P. intermedia, P. jepsonii, P. johnstonii, P. lasiodonta, P. litoralis, P. macounii, P. millefolia, P. modesta, P. morefieldii, P. multijuga, P. multisecta, P. nana, P. newberryi, P. nivea, P. norvegica, P. ovina, P. paucijuga, P. pedersenii, P. pensylvanica, P. pseudosericea, P. pulchella, P. pulcherrima, P. recta, P. reptans, P. rhyolitica, P. rimicola, P. rivalis, P. robbinsiana, P. rubella, P. rubricaulis, P. sanguinea, P. saximontana, P. sierrae-blancae, P. simplex, P. sterilis, P. stipularis, P. subgorodkovii, P. subjuga, P. subvahliana, P. subviscosa, P. supina, P. thurberi, P. thuringiaca, P. tikhomirovii, P. townsendii, P. uliginosa, P. uschakovii, P. vahliana, P. verna, P. versicolor, P. villosa, P. villosula, P. vulcanicola, P. wheeleri
Synonyms P. leucophylla, P. argyrea, P. effusa var. argyrea, P. hippiana var. argyrea, P. hippiana var. diffusa, P. propinqua P. diversifolia var. madsenii, P. plattensis var. pedicillata
Name authority Lehmann: Nov. Stirp. Pug. 2: 7. (1830) — not Pallas 1773 Nuttall: in J. Torrey and A. Gray, Fl. N. Amer. 1: 439. (1840)
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