Potentilla hippiana
Potentilla norvegica
Hipp's cinquefoil, horse cinquefoil, potentille de Hipp, woolly cinquefoil
Norwegian cinquefoil, Norwegian or rough cinquefoil, potentille de norvège, rough cinquefoil
(0.3–)2–4(–5) dm, lengths (1–)2–4 times basal leaves.
ascending to erect, (0.5–)2–5(–9) dm, hairs at base ± stiff, tubercle-based, glands absent or sparse, inconspicuous.
ternate, rarely palmate, 3–15(–20) cm;
petiole 1–6(–10) cm, long hairs sparse to abundant, spreading to ascending, 1–2.5(–3) mm, usually ± stiff, ± crisped hairs absent or sparse to common, glands absent or sparse, inconspicuous;
leaflets 3(–5), at tip of leaf axis, separate to ± overlapping, largest ones broadly oblanceolate or elliptic to obovate, 1–6(–10) × 0.7–4(–5) cm, distal (1/2–)2/3–3/4+ of margin usually ± evenly incised 1/4–1/3 to midvein, teeth (3–)4–8(–15) per side, surfaces sparsely to moderately hairy, sometimes glabrate or abundantly hairy, glands mostly absent.
pinnate to subpinnate, (3–)5–15(–25) cm;
petiole 1–10(–15) cm, long hairs abundant to dense, tightly appressed, 1–2.5 mm, stiff, short hairs absent or obscured, crisped-cottony hairs absent or sparse to common, glands absent or obscured;
leaflets not conduplicate, lateral ones evenly (to unevenly in argyrea phase) paired, (2–)3–6(–7) per side on distal (1/6–)1/5–1/2 of leaf axis, distal pairs ± decurrent, often confluent with terminal leaflet, larger leaflets oblanceolate or narrowly obovate to oblong, 1–5(–6) × 0.3–1.5 cm, distal (2/3–)3/4 to whole margin incised 1/2 or less (rarely more) to midvein, teeth (5–)7–12(–18) per side, 1–4 mm, surfaces ± to strongly dissimilar, abaxial white, long hairs abundant (mostly on, but not limited to, veins), 1–2 mm, stiff, short hairs absent or obscured, crisped-cottony hairs abundant to dense, glands absent or obscured, adaxial green to grayish, long hairs sparse to common, rarely absent, short or crisped to, sometimes, cottony hairs absent or sparse to common, glands sparse.
1–2(–3).
10–30-flowered.
5–40+-flowered.
0.3–3(–5) cm.
(0.2–)0.5–2.5(–3) cm.
epicalyx bractlets narrowly elliptic to lanceolate, rarely linear, 2–5 × 0.5–1.5 mm, (1/2–)2/3 as long as sepals, abaxial vestiture similar to or ± sparser than sepals, usually not glabrescent, straight hairs ± abundant, crisped to sometimes ± cottony hairs absent or sparse to abundant;
hypanthium 3–7 mm diam.;
sepals 4–5.5(–6.5) mm, apex acute to acuminate;
petals 4–8 × 4–7 mm;
filaments 0.5–2.5 mm, anthers 0.6–1.1 mm;
carpels (5–)10–30, styles 1.7–2.5 mm.
epicalyx bractlets ± elliptic to narrowly ovate, (3–)4–8(–13) × 1.5–3(–5) mm;
hypanthium 4–7 mm diam.;
sepals 5–8 mm, apex acute to obtuse;
petals yellow, broadly obovate, (2–)3–5 × 2–4 mm;
stamens 15 or 20, filaments 0.7–2 mm, anthers 0.3–0.5 mm;
carpels 60–150, styles 0.7–0.8 mm.
1.4–1.8 mm, smooth to faintly rugose.
tan to brown, 0.8–1.3 mm, usually strongly rugose, without a corky protuberance.
= 42, 70, 77, 84, 98.
= 42, 56, 63, 70.
Potentilla hippiana
Potentilla norvegica
Potentilla hippiana is most common and distinctive in the Colorado Plateau and southern Rocky Mountains, with outliers in the mountains of Arizona, Nevada, and New Mexico. Occurrences are more scattered in the northern Rocky Mountains and Great Plains, where intergradation with P. effusa is common. Among these intermediates are the argyrea phase and the single report from Idaho (B. C. Johnston 1980). Potentilla hippiana is probably adventive in Nova Scotia, eastern Ontario, Quebec, and Michigan, and possibly the Northwest Territories. The reported occurrence in central Alaska (E. Hultén 1968) is based on a specimen of P. pensylvanica (CAN).
Significant variation exists throughout the range of Potentilla hippiana, including dwarfism, leaf division, leaflet dissection, adaxial leaflet vestiture, and pedicel length. Some of this variation might merit taxonomic recognition upon further analysis, especially if correlated with ploidy level (6x through 12x). At present, the differences used to distinguish var. argyrea are not found to be sufficiently correlated with distribution to merit formal taxonomic recognition.
Complicating the infraspecific variation is a tendency for Potentilla hippiana to intergrade with other species, in particular P. effusa. Beyond sect. Leucophyllae, the frequency of hybrids with P. pulcherrima blurs the distinction between the two species; P. gracilis var. hippianoides S. L. Welsh & N. D. Atwood is probably one of the resultant intermediates.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Potentilla norvegica is considered native in both North America and Eurasia, with the American race occasionally recognized as subsp. hirsuta for the stiff hairs on the stems, petioles, and pedicels of most plants in the flora area. This feature and other purported differences between the two races, in addition to being relatively subtle and inconstant, can be found in both America and Eurasia, though undoubtedly at least in part as introductions. Some populations in eastern Canada with glabrous stems have been distinguished as var. labradorica (for example, M. L. Fernald 1950), but such plants typically are intermixed with hairy individuals. The Löves (Á Löve 1954; Á Löve and D. Löve 1966) have argued that all three variants should be treated as distinct species due to chromosomal differences (hirsuta 2n = 56; labradorica 2n = 42; norvegica in the narrow sense 2n = 70) and obligate apomixis. Further research is needed to determine the taxonomic validity and rank of these expressions. If treated as species, P. flexuosa antedates P. labradorica (as noted by J. Soják 1969), while P. monspeliensis, although commonly used for the American race, is based on a European type (as summarized by A. Kurtto et al. in J. Jalas et al. 1972+, vol. 13).
An even greater challenge is determining native versus introduced ranges, especially given the likelihood that both native and Eurasian populations are widespread in North America. Achenes are produced prolifically and easily dispersed, to the extent that Potentilla norvegica is a contaminant in clover and hay fields and considered a weed at least in Canada (P. A. Werner and J. D. Soule 1976).
(Discussion copyrighted by Flora of North America; reprinted with permission.)
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