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Hipp's cinquefoil, horse cinquefoil, potentille de Hipp, woolly cinquefoil

modest cinquefoil

Caudex branches

usually not sheathed with marcescent whole leaves.

Stems

(0.3–)2–4(–5) dm, lengths (1–)2–4 times basal leaves.

ascending to nearly erect, (0.3–)0.5–1.5(–2.5) dm.

Basal leaves

pinnate to subpinnate, (3–)5–15(–25) cm;

petiole 1–10(–15) cm, long hairs abundant to dense, tightly appressed, 1–2.5 mm, stiff, short hairs absent or obscured, crisped-cottony hairs absent or sparse to common, glands absent or obscured;

leaflets not conduplicate, lateral ones evenly (to unevenly in argyrea phase) paired, (2–)3–6(–7) per side on distal (1/6–)1/5–1/2 of leaf axis, distal pairs ± decurrent, often confluent with terminal leaflet, larger leaflets oblanceolate or narrowly obovate to oblong, 1–5(–6) × 0.3–1.5 cm, distal (2/3–)3/4 to whole margin incised 1/2 or less (rarely more) to midvein, teeth (5–)7–12(–18) per side, 1–4 mm, surfaces ± to strongly dissimilar, abaxial white, long hairs abundant (mostly on, but not limited to, veins), 1–2 mm, stiff, short hairs absent or obscured, crisped-cottony hairs abundant to dense, glands absent or obscured, adaxial green to grayish, long hairs sparse to common, rarely absent, short or crisped to, sometimes, cottony hairs absent or sparse to common, glands sparse.

usually palmate, rarely also ternate on same plant or subpalmate, 2–6(–8) cm;

petiole 0.5–3.5(–5) cm, long hairs common to abundant, ± ascending to almost spreading, 1–2 mm, weak to ± stiff, scarcely to ± verrucose, short and/or crisped hairs common to abundant, cottony hairs absent, glands sparse to common;

leaflets (3–)5, proximalmost separated by 0–1 mm, central obovate to oblanceolate-elliptic, 0.5–2(–2.5) × 0.5–1(–1.5) cm, petiolules 1 mm, distal 3/4 to nearly whole margin incised 1/2–3/4 to midvein, teeth 2–5 per side, (1–)2–5 mm, apical tufts 0.5–1 mm, abaxial surfaces white, long hairs common to abundant, cottony-crisped hairs dense, short hairs and glands absent or obscured, adaxial grayish green, long hairs common to abundant, 0.5–1.5 mm, ± stiff, rarely soft, short and/or crisped hairs sparse to common, cottony hairs absent, glands sparse to common.

Cauline leaves

1–2(–3).

1–2.

Inflorescences

10–30-flowered.

(1–)3–10-flowered, congested or ± elongating in fruit, branch angle 20–35°.

Pedicels

0.3–3(–5) cm.

0.2–0.5 cm, proximal to 1.2 cm.

Flowers

epicalyx bractlets narrowly elliptic to lanceolate, rarely linear, 2–5 × 0.5–1.5 mm, (1/2–)2/3 as long as sepals, abaxial vestiture similar to or ± sparser than sepals, usually not glabrescent, straight hairs ± abundant, crisped to sometimes ± cottony hairs absent or sparse to abundant;

hypanthium 3–7 mm diam.;

sepals 4–5.5(–6.5) mm, apex acute to acuminate;

petals 4–8 × 4–7 mm;

filaments 0.5–2.5 mm, anthers 0.6–1.1 mm;

carpels (5–)10–30, styles 1.7–2.5 mm.

epicalyx bractlets narrowly elliptic, 1.5–3 × 0.5–1 mm;

hypanthium 3–4 mm diam.;

sepals 2.5–4 mm, apex ± acute, glands abundant, usually not obscured;

petals yellow, ± overlapping, 3.5–5 × 3–4 mm, slightly longer than sepals;

filaments 1 mm, anthers 0.3–0.7 mm;

carpels 20–40, styles 1 mm.

Achenes

1.4–1.8 mm, smooth to faintly rugose.

1 mm.

2n

= 42, 70, 77, 84, 98.

Potentilla hippiana

Potentilla modesta

Phenology Flowering summer. Flowering summer.
Habitat Dry grasslands and meadows, in aspen and conifer woodlands or alpine tundra, disturbed sites Alpine tundra, fellfields, talus slopes, cliffs, usually on limestone
Elevation 500–3400 m (1600–11200 ft) 2500–3900 m (8200–12800 ft)
Distribution
from FNA
AZ; CO; ID; MI; MN; MT; ND; NE; NM; NV; SD; UT; WY; AB; BC; MB; NS; NT; ON; QC; SK
[WildflowerSearch map]
[BONAP county map]
from FNA
CO; ID; MT; UT; WY
Discussion

Potentilla hippiana is most common and distinctive in the Colorado Plateau and southern Rocky Mountains, with outliers in the mountains of Arizona, Nevada, and New Mexico. Occurrences are more scattered in the northern Rocky Mountains and Great Plains, where intergradation with P. effusa is common. Among these intermediates are the argyrea phase and the single report from Idaho (B. C. Johnston 1980). Potentilla hippiana is probably adventive in Nova Scotia, eastern Ontario, Quebec, and Michigan, and possibly the Northwest Territories. The reported occurrence in central Alaska (E. Hultén 1968) is based on a specimen of P. pensylvanica (CAN).

Significant variation exists throughout the range of Potentilla hippiana, including dwarfism, leaf division, leaflet dissection, adaxial leaflet vestiture, and pedicel length. Some of this variation might merit taxonomic recognition upon further analysis, especially if correlated with ploidy level (6x through 12x). At present, the differences used to distinguish var. argyrea are not found to be sufficiently correlated with distribution to merit formal taxonomic recognition.

Complicating the infraspecific variation is a tendency for Potentilla hippiana to intergrade with other species, in particular P. effusa. Beyond sect. Leucophyllae, the frequency of hybrids with P. pulcherrima blurs the distinction between the two species; P. gracilis var. hippianoides S. L. Welsh & N. D. Atwood is probably one of the resultant intermediates.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Potentilla modesta is the dominant component of what has previously been called P. rubricaulis in the Intermountain Region (for example, N. H. Holmgren 1997b, including illustration). This and other species of alpine Potentilla often grow in mixed populations, which contributes to confusion and mistaken identifications. In general, P. modesta has a more compact inflorescence and more consistently palmate basal leaves than sympatric members of the section. Intergrades with P. jepsonii (sect. Pensylvanicae) can also be problematic, though leaves of the latter are generally subpinnate and abaxially grayish rather than white.

The epithet modesta is misapplied in the combination Potentilla concinna var. modesta, which S. L. Welsh et al. (1993) used for plants mostly placed here in P. concinna var. divisa (sect. Concinnae).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 9, p. 162. FNA vol. 9, p. 209.
Parent taxa Rosaceae > subfam. Rosoideae > tribe Potentilleae > Potentilla > sect. Leucophyllae Rosaceae > subfam. Rosoideae > tribe Potentilleae > Potentilla > sect. Rubricaules
Sibling taxa
P. albiflora, P. ambigens, P. anachoretica, P. angelliae, P. anglica, P. anserina, P. arenosa, P. argentea, P. arizonica, P. basaltica, P. bicrenata, P. biennis, P. biflora, P. bimundorum, P. bipinnatifida, P. brevifolia, P. breweri, P. bruceae, P. brunnescens, P. canadensis, P. concinna, P. cottamii, P. crantzii, P. crebridens, P. crinita, P. cristae, P. demotica, P. drummondii, P. effusa, P. elegans, P. erecta, P. flabellifolia, P. fragiformis, P. furcata, P. glaucophylla, P. gracilis, P. grayi, P. hickmanii, P. holmgrenii, P. hookeriana, P. hyparctica, P. inclinata, P. intermedia, P. jepsonii, P. johnstonii, P. lasiodonta, P. litoralis, P. macounii, P. millefolia, P. modesta, P. morefieldii, P. multijuga, P. multisecta, P. nana, P. newberryi, P. nivea, P. norvegica, P. ovina, P. paucijuga, P. pedersenii, P. pensylvanica, P. plattensis, P. pseudosericea, P. pulchella, P. pulcherrima, P. recta, P. reptans, P. rhyolitica, P. rimicola, P. rivalis, P. robbinsiana, P. rubella, P. rubricaulis, P. sanguinea, P. saximontana, P. sierrae-blancae, P. simplex, P. sterilis, P. stipularis, P. subgorodkovii, P. subjuga, P. subvahliana, P. subviscosa, P. supina, P. thurberi, P. thuringiaca, P. tikhomirovii, P. townsendii, P. uliginosa, P. uschakovii, P. vahliana, P. verna, P. versicolor, P. villosa, P. villosula, P. vulcanicola, P. wheeleri
P. albiflora, P. ambigens, P. anachoretica, P. angelliae, P. anglica, P. anserina, P. arenosa, P. argentea, P. arizonica, P. basaltica, P. bicrenata, P. biennis, P. biflora, P. bimundorum, P. bipinnatifida, P. brevifolia, P. breweri, P. bruceae, P. brunnescens, P. canadensis, P. concinna, P. cottamii, P. crantzii, P. crebridens, P. crinita, P. cristae, P. demotica, P. drummondii, P. effusa, P. elegans, P. erecta, P. flabellifolia, P. fragiformis, P. furcata, P. glaucophylla, P. gracilis, P. grayi, P. hickmanii, P. hippiana, P. holmgrenii, P. hookeriana, P. hyparctica, P. inclinata, P. intermedia, P. jepsonii, P. johnstonii, P. lasiodonta, P. litoralis, P. macounii, P. millefolia, P. morefieldii, P. multijuga, P. multisecta, P. nana, P. newberryi, P. nivea, P. norvegica, P. ovina, P. paucijuga, P. pedersenii, P. pensylvanica, P. plattensis, P. pseudosericea, P. pulchella, P. pulcherrima, P. recta, P. reptans, P. rhyolitica, P. rimicola, P. rivalis, P. robbinsiana, P. rubella, P. rubricaulis, P. sanguinea, P. saximontana, P. sierrae-blancae, P. simplex, P. sterilis, P. stipularis, P. subgorodkovii, P. subjuga, P. subvahliana, P. subviscosa, P. supina, P. thurberi, P. thuringiaca, P. tikhomirovii, P. townsendii, P. uliginosa, P. uschakovii, P. vahliana, P. verna, P. versicolor, P. villosa, P. villosula, P. vulcanicola, P. wheeleri
Synonyms P. leucophylla, P. argyrea, P. effusa var. argyrea, P. hippiana var. argyrea, P. hippiana var. diffusa, P. propinqua P. concinna var. modesta
Name authority Lehmann: Nov. Stirp. Pug. 2: 7. (1830) — not Pallas 1773 Rydberg: in N. L. Britton et al., N. Amer. Fl. 22: 331. (1908)
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