Potentilla hippiana |
Potentilla gracilis |
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Hipp's cinquefoil, horse cinquefoil, potentille de Hipp, woolly cinquefoil |
graceful cinquefoil, northwest cinquefoil, potentille grêle, slender cinquefoil |
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Glands | usually absent or inconspicuous, uncolored. |
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Stems | (0.3–)2–4(–5) dm, lengths (1–)2–4 times basal leaves. |
ascending to erect, (0.5–)2–7(–12) dm. |
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Basal leaves | pinnate to subpinnate, (3–)5–15(–25) cm; petiole 1–10(–15) cm, long hairs abundant to dense, tightly appressed, 1–2.5 mm, stiff, short hairs absent or obscured, crisped-cottony hairs absent or sparse to common, glands absent or obscured; leaflets not conduplicate, lateral ones evenly (to unevenly in argyrea phase) paired, (2–)3–6(–7) per side on distal (1/6–)1/5–1/2 of leaf axis, distal pairs ± decurrent, often confluent with terminal leaflet, larger leaflets oblanceolate or narrowly obovate to oblong, 1–5(–6) × 0.3–1.5 cm, distal (2/3–)3/4 to whole margin incised 1/2 or less (rarely more) to midvein, teeth (5–)7–12(–18) per side, 1–4 mm, surfaces ± to strongly dissimilar, abaxial white, long hairs abundant (mostly on, but not limited to, veins), 1–2 mm, stiff, short hairs absent or obscured, crisped-cottony hairs abundant to dense, glands absent or obscured, adaxial green to grayish, long hairs sparse to common, rarely absent, short or crisped to, sometimes, cottony hairs absent or sparse to common, glands sparse. |
usually not in ranks, palmate, rarely subpalmate, 4–40(–55) cm; petiole (2–)3–25(–45) cm, long hairs sparse to dense, appressed to spreading, 0.5–2(–3) mm, weak to stiff, short hairs absent, crisped hairs usually absent, cottony hairs absent or sparse to abundant (var. owyheensis), glands absent or sparse; leaflets 5–9, on tip, rarely less than distal 1/10, of leaf axis, separate to overlapping, largest ones oblanceolate to elliptic or obovate, (1.5–)2–9(–11) × (0.7–)1.5–4 cm, margins flat to revolute, distal (2/3–)3/4 to nearly whole length evenly to unevenly incised 1/4–3/4+ to midvein, undivided medial blade 2–20 mm wide, teeth (4–)6–10(–11) per side (sometimes secondarily toothed), linear to broadly lanceolate, 2–20(–35) mm, surfaces ± similar to strongly dissimilar, abaxial green or pale green to white, usually not glaucous, long hairs sparse to abundant (sometimes mostly on veins), short-crisped hairs absent or sparse to abundant, cottony hairs absent or sparse to dense, glands usually absent or obscured to sparse, sometimes common, adaxial dark green to grayish, long hairs sparse to ± abundant, rarely absent or dense, short-crisped hairs absent or sparse, cottony hairs absent or sparse to abundant (var. owyheensis), glands usually absent or sparse to sometimes common. |
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Cauline leaves | 1–2(–3). |
1–4(–5). |
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Inflorescences | 10–30-flowered. |
(4–)10–50(–60)-flowered. |
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Pedicels | 0.3–3(–5) cm. |
0.3–3 cm. |
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Flowers | epicalyx bractlets narrowly elliptic to lanceolate, rarely linear, 2–5 × 0.5–1.5 mm, (1/2–)2/3 as long as sepals, abaxial vestiture similar to or ± sparser than sepals, usually not glabrescent, straight hairs ± abundant, crisped to sometimes ± cottony hairs absent or sparse to abundant; hypanthium 3–7 mm diam.; sepals 4–5.5(–6.5) mm, apex acute to acuminate; petals 4–8 × 4–7 mm; filaments 0.5–2.5 mm, anthers 0.6–1.1 mm; carpels (5–)10–30, styles 1.7–2.5 mm. |
epicalyx bractlets linear to narrowly lanceolate or elliptic, 2–6(–8) × 0.5–1.5(–2.5) mm, hairs sparse to abundant, rarely dense, ± appressed to ascending, rarely spreading, glands usually absent or inconspicuous; hypanthium 3.5–6 mm diam.; sepals 4–8(–10) mm, apex acute to long acuminate; petals (3–)4–10(–11) × (3–)4–10(–12) mm; filaments (1–)1.5–2.5(–3.5) mm, anthers 0.6–1.2(–1.6) mm; carpels (15–)20–50, styles ± tapered, papillate-swollen proximally, (1–)1.5–2.5(–3) mm. |
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Achenes | 1.4–1.8 mm, smooth to faintly rugose. |
1–1.8 mm. |
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2n | = 42, 70, 77, 84, 98. |
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Potentilla hippiana |
Potentilla gracilis |
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Phenology | Flowering summer. | |||||||||||||||||
Habitat | Dry grasslands and meadows, in aspen and conifer woodlands or alpine tundra, disturbed sites | |||||||||||||||||
Elevation | 500–3400 m (1600–11200 ft) | |||||||||||||||||
Distribution |
AZ; CO; ID; MI; MN; MT; ND; NE; NM; NV; SD; UT; WY; AB; BC; MB; NS; NT; ON; QC; SK
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AK; AZ; CA; CO; ID; MI; MN; MT; ND; NV; OR; SD; UT; WA; WI; WY; AB; BC; MB; NT; ON; PE; QC; SK; YT
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Discussion | Potentilla hippiana is most common and distinctive in the Colorado Plateau and southern Rocky Mountains, with outliers in the mountains of Arizona, Nevada, and New Mexico. Occurrences are more scattered in the northern Rocky Mountains and Great Plains, where intergradation with P. effusa is common. Among these intermediates are the argyrea phase and the single report from Idaho (B. C. Johnston 1980). Potentilla hippiana is probably adventive in Nova Scotia, eastern Ontario, Quebec, and Michigan, and possibly the Northwest Territories. The reported occurrence in central Alaska (E. Hultén 1968) is based on a specimen of P. pensylvanica (CAN). Significant variation exists throughout the range of Potentilla hippiana, including dwarfism, leaf division, leaflet dissection, adaxial leaflet vestiture, and pedicel length. Some of this variation might merit taxonomic recognition upon further analysis, especially if correlated with ploidy level (6x through 12x). At present, the differences used to distinguish var. argyrea are not found to be sufficiently correlated with distribution to merit formal taxonomic recognition. Complicating the infraspecific variation is a tendency for Potentilla hippiana to intergrade with other species, in particular P. effusa. Beyond sect. Leucophyllae, the frequency of hybrids with P. pulcherrima blurs the distinction between the two species; P. gracilis var. hippianoides S. L. Welsh & N. D. Atwood is probably one of the resultant intermediates. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Varieties 5 (5 in the flora). Because variation within Potentilla gracilis and other members of sect. Graciles is still far from resolved, taxonomic recognition is given to only the most distinctive extremes and that mostly at the varietal level. The goal is to call attention to areas that need study rather than to provide a full resolution, and even this leaves out some large, widespread populations in both keys and descriptions. The taxonomic placement of some available names, such as P. glomerata A. Nelson, P. longipedunculata Rydberg, and P. macropetala Rydberg, likewise remains unresolved. The vouchers for the report by M. L. Fernald (1950) of Potentilla gracilis and P. pulcherrima from New Hampshire are presumably specimens collected in Coos County by A. S. Pease (NEBC). The three collections differ from one another, corresponding most closely to var. fastigiata, var. flabelliformis, and P. pulcherrima. The presence of such diversity in the same meadow, all as disjunct occurrences, suggests an artificial, non-persisting introduction. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 9, p. 162. | FNA vol. 9, p. 153. | ||||||||||||||||
Parent taxa | Rosaceae > subfam. Rosoideae > tribe Potentilleae > Potentilla > sect. Leucophyllae | Rosaceae > subfam. Rosoideae > tribe Potentilleae > Potentilla > sect. Graciles | ||||||||||||||||
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Subordinate taxa | ||||||||||||||||||
Synonyms | P. leucophylla, P. argyrea, P. effusa var. argyrea, P. hippiana var. argyrea, P. hippiana var. diffusa, P. propinqua | |||||||||||||||||
Name authority | Lehmann: Nov. Stirp. Pug. 2: 7. (1830) — not Pallas 1773 | Douglas ex Hooker: Bot. Mag. 57: plate 2984. (1830) | ||||||||||||||||
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