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Hipp's cinquefoil, horse cinquefoil, potentille de Hipp, woolly cinquefoil

erect cinquefoil, tormentil

Stems

(0.3–)2–4(–5) dm, lengths (1–)2–4 times basal leaves.

mostly ascending to erect, not flagelliform, openly branched, not rooting at nodes, (0.5–)1–5(–7) dm.

Basal leaves

pinnate to subpinnate, (3–)5–15(–25) cm;

petiole 1–10(–15) cm, long hairs abundant to dense, tightly appressed, 1–2.5 mm, stiff, short hairs absent or obscured, crisped-cottony hairs absent or sparse to common, glands absent or obscured;

leaflets not conduplicate, lateral ones evenly (to unevenly in argyrea phase) paired, (2–)3–6(–7) per side on distal (1/6–)1/5–1/2 of leaf axis, distal pairs ± decurrent, often confluent with terminal leaflet, larger leaflets oblanceolate or narrowly obovate to oblong, 1–5(–6) × 0.3–1.5 cm, distal (2/3–)3/4 to whole margin incised 1/2 or less (rarely more) to midvein, teeth (5–)7–12(–18) per side, 1–4 mm, surfaces ± to strongly dissimilar, abaxial white, long hairs abundant (mostly on, but not limited to, veins), 1–2 mm, stiff, short hairs absent or obscured, crisped-cottony hairs abundant to dense, glands absent or obscured, adaxial green to grayish, long hairs sparse to common, rarely absent, short or crisped to, sometimes, cottony hairs absent or sparse to common, glands sparse.

ephemeral, usually ternate, 2–15 cm;

petiole (1–)3–13 cm, long hairs sparse to common, appressed, 0.5–1 mm, stiff, glands absent;

leaflets 3(–5), central ± obovate to cuneate, (0.5–)1–2 × (0.3–)1–1.5 cm, distal 1/2–2/3 of margin incised ± 1/3 to midvein, teeth 2–4 per side, surfaces similar, green (abaxial sometimes paler), glabrate or sparsely to moderately hairy.

Cauline leaves

1–2(–3).

(0–)1–3(–7) proximal to 1st flowering node, expanded at anthesis, usually ternate, rarely palmate (or appearing so because of leafletlike stipules), (0.5–)1–3 cm;

petiole usually 0 cm;

leaflets 3(–5), usually much narrower than those of basal leaves, narrowly cuneate to oblanceolate, (0.5–)1–3 × (0.2–)0.5–1 cm, distal 1/3–2/3 of margin incised ± 1/2 to midvein, teeth 2–4(–6) per side, apex usually obtuse to acute.

Inflorescences

10–30-flowered.

3–30-flowered, cymose, open.

Pedicels

0.3–3(–5) cm.

(1–)2–3(–5) cm.

Flowers

epicalyx bractlets narrowly elliptic to lanceolate, rarely linear, 2–5 × 0.5–1.5 mm, (1/2–)2/3 as long as sepals, abaxial vestiture similar to or ± sparser than sepals, usually not glabrescent, straight hairs ± abundant, crisped to sometimes ± cottony hairs absent or sparse to abundant;

hypanthium 3–7 mm diam.;

sepals 4–5.5(–6.5) mm, apex acute to acuminate;

petals 4–8 × 4–7 mm;

filaments 0.5–2.5 mm, anthers 0.6–1.1 mm;

carpels (5–)10–30, styles 1.7–2.5 mm.

4(–5)-merous;

epicalyx bractlets ovate to narrowly elliptic, 1.5–4 × 0.5–1 mm, much smaller than to sometimes equal to sepals;

hypanthium 1.5–2.5 mm diam.;

sepals (2–)3–5 mm, apex broadly acute;

petals (2–)4–6 × (1.5–)3–6 mm, apex ± retuse;

stamens 15–20, filaments 2–3.8 mm, anthers 0.4–0.6 mm;

carpels 4–8(–20), styles 0.9–1.4 mm.

Achenes

1.4–1.8 mm, smooth to faintly rugose.

1.2–2 mm, rugose.

Rootstocks

erect to horizontal, irregularly thickened, 1–8 cm.

2n

= 42, 70, 77, 84, 98.

= 28 (Eurasia).

Potentilla hippiana

Potentilla erecta

Phenology Flowering summer. Flowering unknown.
Habitat Dry grasslands and meadows, in aspen and conifer woodlands or alpine tundra, disturbed sites Moist, mossy flats and slopes, acidic soil
Elevation 500–3400 m (1600–11200 ft) 0–100 m (0–300 ft)
Distribution
from FNA
AZ; CO; ID; MI; MN; MT; ND; NE; NM; NV; SD; UT; WY; AB; BC; MB; NS; NT; ON; QC; SK
[WildflowerSearch map]
[BONAP county map]
from FNA
MA; NS; Europe; w Asia; n Africa; Atlantic Islands (Azores) [Introduced in North America]
[BONAP county map]
Discussion

Potentilla hippiana is most common and distinctive in the Colorado Plateau and southern Rocky Mountains, with outliers in the mountains of Arizona, Nevada, and New Mexico. Occurrences are more scattered in the northern Rocky Mountains and Great Plains, where intergradation with P. effusa is common. Among these intermediates are the argyrea phase and the single report from Idaho (B. C. Johnston 1980). Potentilla hippiana is probably adventive in Nova Scotia, eastern Ontario, Quebec, and Michigan, and possibly the Northwest Territories. The reported occurrence in central Alaska (E. Hultén 1968) is based on a specimen of P. pensylvanica (CAN).

Significant variation exists throughout the range of Potentilla hippiana, including dwarfism, leaf division, leaflet dissection, adaxial leaflet vestiture, and pedicel length. Some of this variation might merit taxonomic recognition upon further analysis, especially if correlated with ploidy level (6x through 12x). At present, the differences used to distinguish var. argyrea are not found to be sufficiently correlated with distribution to merit formal taxonomic recognition.

Complicating the infraspecific variation is a tendency for Potentilla hippiana to intergrade with other species, in particular P. effusa. Beyond sect. Leucophyllae, the frequency of hybrids with P. pulcherrima blurs the distinction between the two species; P. gracilis var. hippianoides S. L. Welsh & N. D. Atwood is probably one of the resultant intermediates.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Although included here, Potentilla erecta is questionably extant as a naturalized species in North America and is not currently known from Newfoundland where historically reported. It is a medicinal plant called tormentil, high in tannins and strongly astringent. A tea made from the tuberous rootstock is widely used in Europe and parts of Asia for diarrhea, dysentery, sore throats, and related problems. A decoction is a mild antibiotic used for cuts and minor infections, especially as an anti-inflammatory for gingivitis and other gum problems. The rootstock is used as a red dye.

Another synonym with a long history of use is Potentilla tormentilla Necker, illegitimate by virtue of being superfluous.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 9, p. 162. FNA vol. 9, p. 136.
Parent taxa Rosaceae > subfam. Rosoideae > tribe Potentilleae > Potentilla > sect. Leucophyllae Rosaceae > subfam. Rosoideae > tribe Potentilleae > Potentilla > sect. Potentilla
Sibling taxa
P. albiflora, P. ambigens, P. anachoretica, P. angelliae, P. anglica, P. anserina, P. arenosa, P. argentea, P. arizonica, P. basaltica, P. bicrenata, P. biennis, P. biflora, P. bimundorum, P. bipinnatifida, P. brevifolia, P. breweri, P. bruceae, P. brunnescens, P. canadensis, P. concinna, P. cottamii, P. crantzii, P. crebridens, P. crinita, P. cristae, P. demotica, P. drummondii, P. effusa, P. elegans, P. erecta, P. flabellifolia, P. fragiformis, P. furcata, P. glaucophylla, P. gracilis, P. grayi, P. hickmanii, P. holmgrenii, P. hookeriana, P. hyparctica, P. inclinata, P. intermedia, P. jepsonii, P. johnstonii, P. lasiodonta, P. litoralis, P. macounii, P. millefolia, P. modesta, P. morefieldii, P. multijuga, P. multisecta, P. nana, P. newberryi, P. nivea, P. norvegica, P. ovina, P. paucijuga, P. pedersenii, P. pensylvanica, P. plattensis, P. pseudosericea, P. pulchella, P. pulcherrima, P. recta, P. reptans, P. rhyolitica, P. rimicola, P. rivalis, P. robbinsiana, P. rubella, P. rubricaulis, P. sanguinea, P. saximontana, P. sierrae-blancae, P. simplex, P. sterilis, P. stipularis, P. subgorodkovii, P. subjuga, P. subvahliana, P. subviscosa, P. supina, P. thurberi, P. thuringiaca, P. tikhomirovii, P. townsendii, P. uliginosa, P. uschakovii, P. vahliana, P. verna, P. versicolor, P. villosa, P. villosula, P. vulcanicola, P. wheeleri
P. albiflora, P. ambigens, P. anachoretica, P. angelliae, P. anglica, P. anserina, P. arenosa, P. argentea, P. arizonica, P. basaltica, P. bicrenata, P. biennis, P. biflora, P. bimundorum, P. bipinnatifida, P. brevifolia, P. breweri, P. bruceae, P. brunnescens, P. canadensis, P. concinna, P. cottamii, P. crantzii, P. crebridens, P. crinita, P. cristae, P. demotica, P. drummondii, P. effusa, P. elegans, P. flabellifolia, P. fragiformis, P. furcata, P. glaucophylla, P. gracilis, P. grayi, P. hickmanii, P. hippiana, P. holmgrenii, P. hookeriana, P. hyparctica, P. inclinata, P. intermedia, P. jepsonii, P. johnstonii, P. lasiodonta, P. litoralis, P. macounii, P. millefolia, P. modesta, P. morefieldii, P. multijuga, P. multisecta, P. nana, P. newberryi, P. nivea, P. norvegica, P. ovina, P. paucijuga, P. pedersenii, P. pensylvanica, P. plattensis, P. pseudosericea, P. pulchella, P. pulcherrima, P. recta, P. reptans, P. rhyolitica, P. rimicola, P. rivalis, P. robbinsiana, P. rubella, P. rubricaulis, P. sanguinea, P. saximontana, P. sierrae-blancae, P. simplex, P. sterilis, P. stipularis, P. subgorodkovii, P. subjuga, P. subvahliana, P. subviscosa, P. supina, P. thurberi, P. thuringiaca, P. tikhomirovii, P. townsendii, P. uliginosa, P. uschakovii, P. vahliana, P. verna, P. versicolor, P. villosa, P. villosula, P. vulcanicola, P. wheeleri
Synonyms P. leucophylla, P. argyrea, P. effusa var. argyrea, P. hippiana var. argyrea, P. hippiana var. diffusa, P. propinqua Tormentilla erecta
Name authority Lehmann: Nov. Stirp. Pug. 2: 7. (1830) — not Pallas 1773 (Linnaeus) Raeuschel: Nomencl. Bot, 152. (1797)
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