Potentilla hippiana
Potentilla drummondii
Hipp's cinquefoil, horse cinquefoil, potentille de Hipp, woolly cinquefoil
Drummond's cinquefoil
usually absent or inconspicuous, rarely conspicuous, uncolored.
(0.3–)2–4(–5) dm, lengths (1–)2–4 times basal leaves.
decumbent to nearly erect, 1.5–4.5(–6) dm.
pinnate to subpinnate, (3–)5–15(–25) cm;
petiole 1–10(–15) cm, long hairs abundant to dense, tightly appressed, 1–2.5 mm, stiff, short hairs absent or obscured, crisped-cottony hairs absent or sparse to common, glands absent or obscured;
leaflets not conduplicate, lateral ones evenly (to unevenly in argyrea phase) paired, (2–)3–6(–7) per side on distal (1/6–)1/5–1/2 of leaf axis, distal pairs ± decurrent, often confluent with terminal leaflet, larger leaflets oblanceolate or narrowly obovate to oblong, 1–5(–6) × 0.3–1.5 cm, distal (2/3–)3/4 to whole margin incised 1/2 or less (rarely more) to midvein, teeth (5–)7–12(–18) per side, 1–4 mm, surfaces ± to strongly dissimilar, abaxial white, long hairs abundant (mostly on, but not limited to, veins), 1–2 mm, stiff, short hairs absent or obscured, crisped-cottony hairs abundant to dense, glands absent or obscured, adaxial green to grayish, long hairs sparse to common, rarely absent, short or crisped to, sometimes, cottony hairs absent or sparse to common, glands sparse.
sometimes 2-ranked, subpinnate to pinnate (proximal leaflets often doubled, distal leaflets confluent and/or decurrent), (4–)10–25 cm;
petiole 1–10(–15) cm, long hairs absent or sparse to abundant, appressed, 1–1.5 mm, usually stiff, short, crisped, and cottony hairs usually absent, glands absent or (regionally) abundant;
leaflets 5–9, on distal 1/10–1/2(–3/4) of leaf axis, separate to ± overlapping, largest ones obovate to cuneate, (1–)2–5 × (0.7–)1–3(–3.5) cm, margins flat, distal 1/2–3/4 unevenly, sometimes evenly, incised ± 1/2 to midvein (often with additional incisions nearly to midvein), undivided medial blade 3–15 mm wide, teeth 3–7 per side (sometimes secondarily toothed), linear to lanceolate, 3–10 mm, surfaces similar, green, not glaucous, long hairs nearly absent or sparse to common (sometimes restricted to abaxial veins), short, crisped, and cottony hairs usually absent, glands usually absent, sometimes regionally abundant.
1–2(–3).
1–3.
10–30-flowered.
3–15-flowered.
0.3–3(–5) cm.
1–3(–6.5) cm.
epicalyx bractlets narrowly elliptic to lanceolate, rarely linear, 2–5 × 0.5–1.5 mm, (1/2–)2/3 as long as sepals, abaxial vestiture similar to or ± sparser than sepals, usually not glabrescent, straight hairs ± abundant, crisped to sometimes ± cottony hairs absent or sparse to abundant;
hypanthium 3–7 mm diam.;
sepals 4–5.5(–6.5) mm, apex acute to acuminate;
petals 4–8 × 4–7 mm;
filaments 0.5–2.5 mm, anthers 0.6–1.1 mm;
carpels (5–)10–30, styles 1.7–2.5 mm.
epicalyx bractlets lanceolate to narrowly elliptic, 3–5 × 1–1.6 mm, hairs usually sparse, appressed to ascending, glands usually absent, sometimes regionally sparse to common;
hypanthium (3–)4–6(–7) mm diam.;
sepals 5–10 mm, apex acute to acuminate;
petals (6–)7–12 × 5–10 mm;
filaments 2–3.5 mm, anthers 0.7–1 mm;
carpels (20–)25–40, styles filiform above papillate-swollen base, 2–3 mm.
1.4–1.8 mm, smooth to faintly rugose.
1.5–2 mm.
= 42, 70, 77, 84, 98.
= 64, 92, 96, 98–108.
Potentilla hippiana
Potentilla drummondii
Potentilla hippiana is most common and distinctive in the Colorado Plateau and southern Rocky Mountains, with outliers in the mountains of Arizona, Nevada, and New Mexico. Occurrences are more scattered in the northern Rocky Mountains and Great Plains, where intergradation with P. effusa is common. Among these intermediates are the argyrea phase and the single report from Idaho (B. C. Johnston 1980). Potentilla hippiana is probably adventive in Nova Scotia, eastern Ontario, Quebec, and Michigan, and possibly the Northwest Territories. The reported occurrence in central Alaska (E. Hultén 1968) is based on a specimen of P. pensylvanica (CAN).
Significant variation exists throughout the range of Potentilla hippiana, including dwarfism, leaf division, leaflet dissection, adaxial leaflet vestiture, and pedicel length. Some of this variation might merit taxonomic recognition upon further analysis, especially if correlated with ploidy level (6x through 12x). At present, the differences used to distinguish var. argyrea are not found to be sufficiently correlated with distribution to merit formal taxonomic recognition.
Complicating the infraspecific variation is a tendency for Potentilla hippiana to intergrade with other species, in particular P. effusa. Beyond sect. Leucophyllae, the frequency of hybrids with P. pulcherrima blurs the distinction between the two species; P. gracilis var. hippianoides S. L. Welsh & N. D. Atwood is probably one of the resultant intermediates.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
The present circumscription of Potentilla drummondii differs from that of B. Ertter (1992, 1993), which encompassed P. breweri and P. bruceae as varieties (see 28. P. bruceae discussion for details). The description above focuses on large, eglandular, subpinnate plants that occur in mountains from the central Sierra Nevada and northern Coast Ranges of California to the Olympic Peninsula of Washington, east to southern Alberta. At least some collections previously identified as P. glaucophylla (as P. diversifolia Lehmann) from the Kenai Peninsula in Alaska can be readily accommodated within this concept, as can collections from the Cabinet Mountains in Montana. The distinction between P. drummondii and subpalmate P. glaucophylla can be vague; in general, the proximalmost leaflets of P. glaucophylla are relatively small and often entire; in P. drummondii proximalmost leaflets are similar in size and dissection to other leaflets.
Pinnate-leaved regional variants merit further attention; in particular the smaller form described as Potentilla cascadensis, and glandular plants in the northern Coast Ranges of California. Relatively small plants forming uniform populations in the southern Sierra Nevada may represent stabilized hybrids with P. breweri.
The inclusion of Potentilla bruceae and P. drummondii in sect. Graciles differs from that of J. Clausen et al. (1940) and B. C. Johnston (1980, 1985), who allied both species with P. breweri in sect. Multijugae. Although subpalmate to subpinnate leaves are anomalous in primarily palmate sect. Graciles, doubled proximal leaflets and an indistinct transition between the rachis and axis of the pinnatisect terminal leaflet in these two species suggest that this state is derived from a palmate progenitor. Habit and petioles also are more compatible with sect. Graciles than sect. Multijugae.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
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