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Hipp's cinquefoil, horse cinquefoil, potentille de Hipp, woolly cinquefoil

alpine cinquefoil, potentille de crantz

Habit Plants tufted to ± matted; caudex branches usually short, stout, sometimes elongate, slender.
Stems

(0.3–)2–4(–5) dm, lengths (1–)2–4 times basal leaves.

ascending to erect, 0.5–2(–3) dm, lengths 1.5–3 times basal leaves.

Basal leaves

pinnate to subpinnate, (3–)5–15(–25) cm;

petiole 1–10(–15) cm, long hairs abundant to dense, tightly appressed, 1–2.5 mm, stiff, short hairs absent or obscured, crisped-cottony hairs absent or sparse to common, glands absent or obscured;

leaflets not conduplicate, lateral ones evenly (to unevenly in argyrea phase) paired, (2–)3–6(–7) per side on distal (1/6–)1/5–1/2 of leaf axis, distal pairs ± decurrent, often confluent with terminal leaflet, larger leaflets oblanceolate or narrowly obovate to oblong, 1–5(–6) × 0.3–1.5 cm, distal (2/3–)3/4 to whole margin incised 1/2 or less (rarely more) to midvein, teeth (5–)7–12(–18) per side, 1–4 mm, surfaces ± to strongly dissimilar, abaxial white, long hairs abundant (mostly on, but not limited to, veins), 1–2 mm, stiff, short hairs absent or obscured, crisped-cottony hairs abundant to dense, glands absent or obscured, adaxial green to grayish, long hairs sparse to common, rarely absent, short or crisped to, sometimes, cottony hairs absent or sparse to common, glands sparse.

2-ranked, usually palmate, sometimes ternate, 2–10(–15) cm;

stipules: apex obtuse to truncate, rarely acute;

petiole 0.5–6.5(–10) cm, long hairs usually ± sparse, usually ± ascending, sometimes spreading or appressed, 1–2.5 mm, weak to ± stiff, glands absent or sparse;

leaflets 3–5, central obovate, 1.5–3(–3.5) × 0.8–1.5(–2) cm, petiolule 0–1 mm, margins flat, not lobed, distal 1/2–2/3 evenly incised 1/3–1/2 to midvein, teeth 3–4(–5) per side, not secondarily toothed, surfaces similar, green, not glaucous, hairs absent or sparse, rarely common, 0.8–1.5 mm, glands absent or sparse to abundant (fewer adaxially).

Cauline leaves

1–2(–3).

Inflorescences

10–30-flowered.

(2–)3–8(–12)-flowered.

Pedicels

0.3–3(–5) cm.

straight, 1–3(–4) cm in flower, to 6 cm in fruit.

Flowers

epicalyx bractlets narrowly elliptic to lanceolate, rarely linear, 2–5 × 0.5–1.5 mm, (1/2–)2/3 as long as sepals, abaxial vestiture similar to or ± sparser than sepals, usually not glabrescent, straight hairs ± abundant, crisped to sometimes ± cottony hairs absent or sparse to abundant;

hypanthium 3–7 mm diam.;

sepals 4–5.5(–6.5) mm, apex acute to acuminate;

petals 4–8 × 4–7 mm;

filaments 0.5–2.5 mm, anthers 0.6–1.1 mm;

carpels (5–)10–30, styles 1.7–2.5 mm.

epicalyx bractlets ovate or narrowly ovate, (1.5–)2.5–4 × (0.8–)1–1.3 mm, margins flat;

hypanthium 3–4 mm diam.;

sepals (3–)4–5(–6) mm, apex broadly acute;

petals yellow, 4–7(–9) × 4–8 mm;

filaments 1.8–2.2 mm, anthers 0.3–0.6 mm;

carpels 30–40, styles columnar-filiform, not papillate-swollen proximally, 1–1.6 mm.

Achenes

1.4–1.8 mm, smooth to faintly rugose.

1.2 mm.

2n

= 42, 70, 77, 84, 98.

= 42; 28, 35, 49 (Europe).

Potentilla hippiana

Potentilla crantzii

Phenology Flowering summer. Flowering spring–summer.
Habitat Dry grasslands and meadows, in aspen and conifer woodlands or alpine tundra, disturbed sites Herb meadows and slopes, usually on calcareous substrates, usually near coast
Elevation 500–3400 m (1600–11200 ft) 0–1700 m (0–5600 ft)
Distribution
from FNA
AZ; CO; ID; MI; MN; MT; ND; NE; NM; NV; SD; UT; WY; AB; BC; MB; NS; NT; ON; QC; SK
[WildflowerSearch map]
[BONAP county map]
from FNA
NL; NU; ON; QC; Greenland; Eurasia [Reportedly introduced in s Australia]
[BONAP county map]
Discussion

Potentilla hippiana is most common and distinctive in the Colorado Plateau and southern Rocky Mountains, with outliers in the mountains of Arizona, Nevada, and New Mexico. Occurrences are more scattered in the northern Rocky Mountains and Great Plains, where intergradation with P. effusa is common. Among these intermediates are the argyrea phase and the single report from Idaho (B. C. Johnston 1980). Potentilla hippiana is probably adventive in Nova Scotia, eastern Ontario, Quebec, and Michigan, and possibly the Northwest Territories. The reported occurrence in central Alaska (E. Hultén 1968) is based on a specimen of P. pensylvanica (CAN).

Significant variation exists throughout the range of Potentilla hippiana, including dwarfism, leaf division, leaflet dissection, adaxial leaflet vestiture, and pedicel length. Some of this variation might merit taxonomic recognition upon further analysis, especially if correlated with ploidy level (6x through 12x). At present, the differences used to distinguish var. argyrea are not found to be sufficiently correlated with distribution to merit formal taxonomic recognition.

Complicating the infraspecific variation is a tendency for Potentilla hippiana to intergrade with other species, in particular P. effusa. Beyond sect. Leucophyllae, the frequency of hybrids with P. pulcherrima blurs the distinction between the two species; P. gracilis var. hippianoides S. L. Welsh & N. D. Atwood is probably one of the resultant intermediates.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Potentilla crantzii is primarily European, not restricted to coasts as in North America. The nomenclatural history is long and complex (A. Kurtto et al. in J. Jalas et al. 1972+, vol. 13), with P. alpestris and P. maculata in greatest historical use. The name P. ×protea Soják is available for presumed hybrids between P. crantzii and P. hyparctica.

The emphasis on two-ranked leaves to distinguish Potentilla crantzii from P. verna follows J. Soják (pers. comm.). Some collections of P. crantzii from Newfoundland approach P. verna in their more elongate caudex branches and shorter stature.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 9, p. 162. FNA vol. 9, p. 190.
Parent taxa Rosaceae > subfam. Rosoideae > tribe Potentilleae > Potentilla > sect. Leucophyllae Rosaceae > subfam. Rosoideae > tribe Potentilleae > Potentilla > sect. Aureae
Sibling taxa
P. albiflora, P. ambigens, P. anachoretica, P. angelliae, P. anglica, P. anserina, P. arenosa, P. argentea, P. arizonica, P. basaltica, P. bicrenata, P. biennis, P. biflora, P. bimundorum, P. bipinnatifida, P. brevifolia, P. breweri, P. bruceae, P. brunnescens, P. canadensis, P. concinna, P. cottamii, P. crantzii, P. crebridens, P. crinita, P. cristae, P. demotica, P. drummondii, P. effusa, P. elegans, P. erecta, P. flabellifolia, P. fragiformis, P. furcata, P. glaucophylla, P. gracilis, P. grayi, P. hickmanii, P. holmgrenii, P. hookeriana, P. hyparctica, P. inclinata, P. intermedia, P. jepsonii, P. johnstonii, P. lasiodonta, P. litoralis, P. macounii, P. millefolia, P. modesta, P. morefieldii, P. multijuga, P. multisecta, P. nana, P. newberryi, P. nivea, P. norvegica, P. ovina, P. paucijuga, P. pedersenii, P. pensylvanica, P. plattensis, P. pseudosericea, P. pulchella, P. pulcherrima, P. recta, P. reptans, P. rhyolitica, P. rimicola, P. rivalis, P. robbinsiana, P. rubella, P. rubricaulis, P. sanguinea, P. saximontana, P. sierrae-blancae, P. simplex, P. sterilis, P. stipularis, P. subgorodkovii, P. subjuga, P. subvahliana, P. subviscosa, P. supina, P. thurberi, P. thuringiaca, P. tikhomirovii, P. townsendii, P. uliginosa, P. uschakovii, P. vahliana, P. verna, P. versicolor, P. villosa, P. villosula, P. vulcanicola, P. wheeleri
P. albiflora, P. ambigens, P. anachoretica, P. angelliae, P. anglica, P. anserina, P. arenosa, P. argentea, P. arizonica, P. basaltica, P. bicrenata, P. biennis, P. biflora, P. bimundorum, P. bipinnatifida, P. brevifolia, P. breweri, P. bruceae, P. brunnescens, P. canadensis, P. concinna, P. cottamii, P. crebridens, P. crinita, P. cristae, P. demotica, P. drummondii, P. effusa, P. elegans, P. erecta, P. flabellifolia, P. fragiformis, P. furcata, P. glaucophylla, P. gracilis, P. grayi, P. hickmanii, P. hippiana, P. holmgrenii, P. hookeriana, P. hyparctica, P. inclinata, P. intermedia, P. jepsonii, P. johnstonii, P. lasiodonta, P. litoralis, P. macounii, P. millefolia, P. modesta, P. morefieldii, P. multijuga, P. multisecta, P. nana, P. newberryi, P. nivea, P. norvegica, P. ovina, P. paucijuga, P. pedersenii, P. pensylvanica, P. plattensis, P. pseudosericea, P. pulchella, P. pulcherrima, P. recta, P. reptans, P. rhyolitica, P. rimicola, P. rivalis, P. robbinsiana, P. rubella, P. rubricaulis, P. sanguinea, P. saximontana, P. sierrae-blancae, P. simplex, P. sterilis, P. stipularis, P. subgorodkovii, P. subjuga, P. subvahliana, P. subviscosa, P. supina, P. thurberi, P. thuringiaca, P. tikhomirovii, P. townsendii, P. uliginosa, P. uschakovii, P. vahliana, P. verna, P. versicolor, P. villosa, P. villosula, P. vulcanicola, P. wheeleri
Synonyms P. leucophylla, P. argyrea, P. effusa var. argyrea, P. hippiana var. argyrea, P. hippiana var. diffusa, P. propinqua Fragaria crantzii, P. alpestris, P. crantzii var. hirta, P. flabellifolia var. hirta, P. langeana, P. maculata
Name authority Lehmann: Nov. Stirp. Pug. 2: 7. (1830) — not Pallas 1773 (Crantz) Beck ex Fritsch: Excursionfl. Oesterreich 295. (1897)
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