Potentilla glaucophylla |
Potentilla tikhomirovii |
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blue-leaf cinquefoil, different-leaf cinquefoil, diverse-leaf cinquefoil, mountain meadow cinquefoil, vari-leaf cinquefoil |
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| Habit | Plants ± tufted. | |||||
| Glands | absent or inconspicuous, uncolored. |
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| Caudex branches | slender to stout, not columnar, not sheathed with marcescent whole leaves. |
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| Stems | mostly ± ascending, 0.5–3(–4.5) dm. |
± ascending, sometimes nearly erect, 0.5–1.5(–3) dm, lengths 2–4 times basal leaves. |
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| Basal leaves | often 2-ranked, palmate, sometimes subpalmate, 2–20 cm; petiole (0.5–)1–8(–14) cm, long hairs often absent, sometimes sparse to abundant, usually appressed, 1–2 mm, weak to ± stiff, short, crisped, and cottony hairs absent, glands absent or sparse; leaflets 5–6(–7), sometimes with 1–2(–4) additional, smaller, entire leaflets, on tip or to distal 1/10(–1/4) of leaf axis, separate, largest ones oblanceolate or cuneate to obovate, 1–4(–6) × 0.5–1.5(–2) cm, margins flat, distal 1/3–1/2(–2/3) ± evenly incised 1/2 or nearly to midvein, undivided medial blade 1.5–7(–10) mm wide, teeth 1–3(–5) per side, broadly lanceolate or linear to narrowly oblong, 1–5(–8) mm, surfaces similar adaxial ± less hairy, ± blue-green, usually glaucous, long hairs sparse to abundant (often restricted to veins, distal teeth, and margins), short, crisped, and cottony hairs absent, glands absent or sparse. |
2–4 cm; petiole 1–2.5 cm, long hairs common to abundant, spreading to ascending, rarely loosely appressed, 1–2 mm, ± stiff, verrucose, short-crisped hairs sparse to abundant, sometimes nearly absent, cottony hairs absent, glands sparse; leaflets ± overlapping, central elliptic to obovate, 1–1.5 × 0.6–1.1 cm, petiolulate, base broadly cuneate, margins flat or slightly revolute, distal ± 2/3 incised ± 1/2 to midvein, teeth (3–)4 per side, ± approximate, surfaces ± dissimilar, abaxial reddish or greenish gray to white, long hairs 1 mm, cottony-crisped hairs common to dense, adaxial green or reddish to gray-green, long hairs sparse to abundant, short-crisped hairs absent or sparse to common. |
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| Cauline leaves | (0–)1–2(–3). |
0–1. |
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| Inflorescences | 2–10(–20)-flowered. |
2–5-flowered. |
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| Pedicels | (0.5–)1–3(–6.5) cm. |
1–2 cm in flower, to 3(–5) cm in fruit. |
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| Flowers | epicalyx bractlets lanceolate to elliptic, 2–5 × 0.8–1.2(–1.6) mm, hairs sparse to common, rarely abundant, ± ascending, glands absent or inconspicuous; hypanthium 3.5–5 mm diam.; sepals (2–)2.5–4.5(–5) mm, apex acute to short acuminate; petals (4–)5–10(–12) × 4–9(–10) mm; filaments 1.5–2.5(–3) mm, anthers 0.4–0.7(–0.9) mm; carpels 25–40, styles filiform above papillate-swollen base, 1.5–2.5(–3) mm. |
epicalyx bractlets narrowly elliptic to narrowly ovate, 4–6 × 1–2.5 mm, (1/2–)2/3 to as wide as sepals, margins revolute, sometimes flat, red glands usually common, conspicuous; hypanthium 3.5–4.5 mm diam.; sepals 5–7 mm, apex subacute; petals 6–8 × (4–)5–8 mm, longer than sepals; filaments 0.7–1.2 mm, anthers 0.4 mm; carpels 40–60, apical hairs absent, styles columnar, strongly papillate-swollen in proximal 1/5–1/3, 0.9–1 mm. |
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| Achenes | 1.2–1.6 mm. |
1.2–1.8 mm. |
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| 2n | = 42, 49 (Russian Far East). |
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Potentilla glaucophylla |
Potentilla tikhomirovii |
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| Phenology | Flowering summer. | |||||
| Habitat | Dry tundra meadows, loamy soil banks, sedge-herb slopes | |||||
| Elevation | 0–1100 m [0–3600 ft] | |||||
| Distribution |
AK; CA; CO; ID; MT; NV; OR; UT; WA; WY; AB; BC; NL; NT; SK; YT
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AK; NT; NU; Greenland; Europe (Russia); Asia |
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| Discussion | Varieties 2 (2 in the flora). Potentilla glaucophylla replaces P. diversifolia; an examination of the lectotype of the latter confirms J. Soják’s (1996) conclusion that P. diversifolia applies to a hybrid involving P. glaucophylla and P. hippiana. Historically, this species was often termed P. dissecta Pursh, a misapplied name that has been rejected (B. Ertter et al. 2008). In addition to the nomenclatural change, the circumscription of Potentilla glaucophylla is here restricted to plants with usually glaucous, blue-green, distally toothed leaflets that are usually glabrate (at least in var. glaucophylla). Plants from the Colorado Plateau and southern Rocky Mountains formerly included in this species and having dark green leaves, more leaflet teeth, and larger anthers are transferred either to P. townsendii or to a currently undescribed entity. The two-ranked leaves of Potentilla glaucophylla suggest a possible connection to P. crantzii in sect. Aureae. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Potentilla tikhomirovii is an assumed hybrid species combining characteristics from P. arenosa subsp. arenosa of sect. Niveae (abaxial leaflet surfaces with crisped-cottony hairs, adaxial surfaces with both long and short hairs, petioles often with both long and short hairs, and flower shape and size) and P. hyparctica of sect. Aureae (crisped-cottony hairs often not fully covering abaxial leaf surfaces, reddish color in most plant parts, sepals and epicalyx bractlets relatively large, broad, less acute, and with many and reddish glands, which are often also on petioles and stipules). Potentilla tikhomirovii has an interrupted range and is not very coherent morphologically. It forms large populations locally, probably by agamic seed propagation, and is a significant part of the Potentilla variation in northern and northeastern Greenland, Ellesmere Island, and parts of the Brooks Range, northern Alaska. Both the distributional and the morphologic patterns indicate that it has evolved multiple times in different regions, such that it is a borderline case for status as an independent species. Its large populations and significant distribution in some regions support recognition, as also does the paucity of obvious back-crosses with the parents. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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| Key |
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| Synonyms | P. diversifolia var. glaucophylla | |||||
| Name authority | Lehmann: Index Seminum (Hamburg) 1836: 7. (1836) | Jurtzev: in A. I. Tolmatchew, Fl. Arct. URSS 9(1): 318. (1984) | ||||
| Source | FNA vol. 9, p. 152. | FNA vol. 9, p. 201. | ||||
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