Potentilla glaucophylla
Potentilla pensylvanica
blue-leaf cinquefoil, different-leaf cinquefoil, diverse-leaf cinquefoil, mountain meadow cinquefoil, vari-leaf cinquefoil
Norwegian cinquefoil, Pennsylvania cinquefoil, pennsylvanian cinquefoil, potentille de pennsylvanie, prairie cinquefoil
absent or inconspicuous, uncolored.
mostly ± ascending, 0.5–3(–4.5) dm.
ascending to erect, (0.3–)1–4(–6) dm.
often 2-ranked, palmate, sometimes subpalmate, 2–20 cm;
petiole (0.5–)1–8(–14) cm, long hairs often absent, sometimes sparse to abundant, usually appressed, 1–2 mm, weak to ± stiff, short, crisped, and cottony hairs absent, glands absent or sparse;
leaflets 5–6(–7), sometimes with 1–2(–4) additional, smaller, entire leaflets, on tip or to distal 1/10(–1/4) of leaf axis, separate, largest ones oblanceolate or cuneate to obovate, 1–4(–6) × 0.5–1.5(–2) cm, margins flat, distal 1/3–1/2(–2/3) ± evenly incised 1/2 or nearly to midvein, undivided medial blade 1.5–7(–10) mm wide, teeth 1–3(–5) per side, broadly lanceolate or linear to narrowly oblong, 1–5(–8) mm, surfaces similar adaxial ± less hairy, ± blue-green, usually glaucous, long hairs sparse to abundant (often restricted to veins, distal teeth, and margins), short, crisped, and cottony hairs absent, glands absent or sparse.
pinnate, (2–)4–20(–25) cm;
petiole 1–8 cm, long hairs sparse to abundant, rarely absent, spreading to ascending, 0.5–3 mm, weak to ± stiff, short hairs abundant to dense, rarely absent, cottony or crisped hairs absent, glands sparse to abundant, often obscured;
leaflets (2–)3–5(–7) per side, on distal 1/2–3/5 of leaf axis, separate to scarcely overlapping, terminal ones narrowly to broadly oblanceolate, (1–)1.5–5(–6) × 0.5–1.5(–2) cm, margins strongly revolute, incised ± 3/4 to midvein, sometimes more or only ± 1/2, undivided medial blade 2–4(–8) mm wide, teeth 4–8(–10) per side, triangular-ovate to linear-elliptic, surfaces usually ± dissimilar, abaxial ± grayish to nearly white, rarely green, long hairs sparse to abundant (mostly on veins), rarely absent, 0.5–1.5 mm, ± weak, short hairs usually abundant to dense, rarely absent, crisped and cottony hairs absent, glands ± abundant, sometimes obscured, adaxial green to gray-green, long hairs sparse to abundant, rarely absent, spreading to ascending, ± 1 mm, short hairs ± abundant, sometimes obscured, rarely absent, cottony and crisped hairs absent, glands ± abundant.
(0–)1–2(–3).
1–4.
2–10(–20)-flowered.
(1–)3–40-flowered, congested or elongating in fruit.
(0.5–)1–3(–6.5) cm.
0.2–1 cm (proximal to 3.5 cm).
epicalyx bractlets lanceolate to elliptic, 2–5 × 0.8–1.2(–1.6) mm, hairs sparse to common, rarely abundant, ± ascending, glands absent or inconspicuous;
hypanthium 3.5–5 mm diam.;
sepals (2–)2.5–4.5(–5) mm, apex acute to short acuminate;
petals (4–)5–10(–12) × 4–9(–10) mm;
filaments 1.5–2.5(–3) mm, anthers 0.4–0.7(–0.9) mm;
carpels 25–40, styles filiform above papillate-swollen base, 1.5–2.5(–3) mm.
epicalyx bractlets usually ± lanceolate-elliptic, rarely ± ovate-elliptic, 3–6(–8) × (1–)1.5–2(–2.5) mm, lengths ± equal to sepals, margins ± revolute;
hypanthium 3–6 mm diam.;
sepals (3–)4–6(–8) mm, apex acute to obtuse, abaxial surfaces: venation indistinct, glands abundant, ± evident;
petals pale to bright yellow, (3–)4–7 × (3–)4–7 mm, lengths ± equal to sepals;
filaments (0.5–)1–2 mm, anthers 0.5–0.8(–1) mm;
carpels 50–100, styles papillate-swollen in proximal ± 1/2, ± 1 mm.
1.2–1.6 mm.
1.2 mm, ± to strongly rugose.
= 28.
Potentilla glaucophylla
Potentilla pensylvanica
Varieties 2 (2 in the flora).
Potentilla glaucophylla replaces P. diversifolia; an examination of the lectotype of the latter confirms J. Soják’s (1996) conclusion that P. diversifolia applies to a hybrid involving P. glaucophylla and P. hippiana. Historically, this species was often termed P. dissecta Pursh, a misapplied name that has been rejected (B. Ertter et al. 2008).
In addition to the nomenclatural change, the circumscription of Potentilla glaucophylla is here restricted to plants with usually glaucous, blue-green, distally toothed leaflets that are usually glabrate (at least in var. glaucophylla). Plants from the Colorado Plateau and southern Rocky Mountains formerly included in this species and having dark green leaves, more leaflet teeth, and larger anthers are transferred either to P. townsendii or to a currently undescribed entity.
The two-ranked leaves of Potentilla glaucophylla suggest a possible connection to P. crantzii in sect. Aureae.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Even in the reduced circumscription used here, Potentilla pensylvanica is exceptionally widespread, occurring in North America from Alaska to Hudson Bay, in the Great Plains and the North American Cordillera to northern Mexico. The species also grows across Siberia (where it has sometimes been called P. sibirica) and as isolated occurrences in the southwestern Alps and Spain (J. Soják 1987c; A. Kurtto et al. in J. Jalas et al. 1972+, vol. 13). One place P. pensylvanica does not occur is in Pennsylvania; the epithet was applied by Linnaeus to cultivated plants originating from Canada. Collections from Quebec are probably introductions (J. Cayouette, pers. comm.). The Oregon citation of P. pensylvanica (D. Mansfield 2000) refers to P. jepsonii.
Significant variation occurs within Potentilla pensylvanica, especially regarding vestiture and plant stature. J. Soják (1987c) has described several varieties, all Eurasian except var. brevipila (long hairs 0.6–1 mm); diminutive specimens from the Brooks Range of northern Alaska, isolated from other known populations of P. pensylvanica, have been described as P. brooksensis. Although much of the ecogeographic variation may merit recognition, at least at varietal level, doing so is postponed pending further analysis and clarification of nomenclature. Among the extremes are plants from the Great Plains, which have abundant long silky hairs overlying and obscuring the velvety layer of short hairs (resembling sympatric P. litoralis), and glabrate (except for glands) populations in Alaska. E. Hultén (1968) erroneously referred to the latter as P. pensylvanica var. glabrata (Lehmann) S. Watson; that name is a synonym of P. ovina var. ovina (sect. Multijugae). Most populations in Alaska also have larger than average flowers (petals to 7 × 7 mm, anthers ca. 1 mm).
Although Potentilla pensylvanica var. strigosa Pursh has been one of the most commonly used varieties in North America, the name has been effectively lectotypified by J. Soják (1987c) on a collection from Siberia. Soják identified this collection as the hybrid between P. pensylvanica and the Asian P. sanguisorba D. F. K. Schlechtendal; var. strigosa accordingly does not occur in North America.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
1. Leaflets incised 1/3–1/2 to midvein, teeth broadly lanceolate; range of species. | var. glaucophylla |
1. Leaflets incised 3/4+ to midvein, teeth linear to narrowly oblong; northern Rocky Mountains. | var. perdissecta |
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