Potentilla glaucophylla |
Potentilla hickmanii |
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blue-leaf cinquefoil, different-leaf cinquefoil, diverse-leaf cinquefoil, mountain meadow cinquefoil, vari-leaf cinquefoil |
Hickman's cinquefoil |
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Habit | Plants ± rosetted; taproots fleshy-thickened. | |||||
Glands | absent or inconspicuous, uncolored. |
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Stems | mostly ± ascending, 0.5–3(–4.5) dm. |
usually prostrate to decumbent, sometimes ± ascending in supporting vegetation, 0.5–2.5 dm, lengths 1–2 times basal leaves. |
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Basal leaves | often 2-ranked, palmate, sometimes subpalmate, 2–20 cm; petiole (0.5–)1–8(–14) cm, long hairs often absent, sometimes sparse to abundant, usually appressed, 1–2 mm, weak to ± stiff, short, crisped, and cottony hairs absent, glands absent or sparse; leaflets 5–6(–7), sometimes with 1–2(–4) additional, smaller, entire leaflets, on tip or to distal 1/10(–1/4) of leaf axis, separate, largest ones oblanceolate or cuneate to obovate, 1–4(–6) × 0.5–1.5(–2) cm, margins flat, distal 1/3–1/2(–2/3) ± evenly incised 1/2 or nearly to midvein, undivided medial blade 1.5–7(–10) mm wide, teeth 1–3(–5) per side, broadly lanceolate or linear to narrowly oblong, 1–5(–8) mm, surfaces similar adaxial ± less hairy, ± blue-green, usually glaucous, long hairs sparse to abundant (often restricted to veins, distal teeth, and margins), short, crisped, and cottony hairs absent, glands absent or sparse. |
pinnate with distal leaflets ± confluent, 3–17 × 1–3.5 cm; petiole 1–5 cm, straight hairs sparse to common, appressed, 0.5–1.5(–2) mm, stiff, cottony hairs absent, glands absent or sparse; primary lateral leaflets 3–6 per side, on distal 1/2–2/3 of leaf axis, ± overlapping to nearly separate, largest ones cuneate to flabellate, 0.5–2 × 0.5–1.5 cm, distal 1/2 to whole margin ± palmately incised 1/2–2/3(+) to midvein, ultimate teeth or segments 2–5, narrowly elliptic to oblanceolate, 2–10(–15) × (1–)1.5–3 mm, apical tufts 0.5 mm, surfaces green, not glaucous, straight hairs sparse to common (often sparser to glabrate adaxially), appressed, 1–1.5 mm, stiff, cottony hairs absent, glands absent or inconspicuous. |
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Cauline leaves | (0–)1–2(–3). |
2. |
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Inflorescences | 2–10(–20)-flowered. |
2–5-flowered, very openly cymose, sometimes racemiform. |
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Pedicels | (0.5–)1–3(–6.5) cm. |
1–2(–4) cm, ± recurved in fruit. |
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Flowers | epicalyx bractlets lanceolate to elliptic, 2–5 × 0.8–1.2(–1.6) mm, hairs sparse to common, rarely abundant, ± ascending, glands absent or inconspicuous; hypanthium 3.5–5 mm diam.; sepals (2–)2.5–4.5(–5) mm, apex acute to short acuminate; petals (4–)5–10(–12) × 4–9(–10) mm; filaments 1.5–2.5(–3) mm, anthers 0.4–0.7(–0.9) mm; carpels 25–40, styles filiform above papillate-swollen base, 1.5–2.5(–3) mm. |
epicalyx bractlets ± elliptic, 3.5–6 × (1–)1.5–2.5 mm; hypanthium 3–5 mm diam.; sepals 3.5–7(–9) mm, apex acute; petals 6–12 × 5–9 mm; filaments (1.5–)2–3(–4) mm, anthers 0.7–1.2 mm; carpels (2–)5–15, styles 2.5–3.5 mm. |
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Achenes | 1.2–1.6 mm. |
2 mm, smooth, ± carunculate. |
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Potentilla glaucophylla |
Potentilla hickmanii |
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Phenology | Flowering spring–early summer. | |||||
Habitat | Vernally saturated coastal meadows, openings in Monterey Pine forests | |||||
Elevation | 30–100 m (100–300 ft) | |||||
Distribution |
AK; CA; CO; ID; MT; NV; OR; UT; WA; WY; AB; BC; NL; NT; SK; YT
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CA |
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Discussion | Varieties 2 (2 in the flora). Potentilla glaucophylla replaces P. diversifolia; an examination of the lectotype of the latter confirms J. Soják’s (1996) conclusion that P. diversifolia applies to a hybrid involving P. glaucophylla and P. hippiana. Historically, this species was often termed P. dissecta Pursh, a misapplied name that has been rejected (B. Ertter et al. 2008). In addition to the nomenclatural change, the circumscription of Potentilla glaucophylla is here restricted to plants with usually glaucous, blue-green, distally toothed leaflets that are usually glabrate (at least in var. glaucophylla). Plants from the Colorado Plateau and southern Rocky Mountains formerly included in this species and having dark green leaves, more leaflet teeth, and larger anthers are transferred either to P. townsendii or to a currently undescribed entity. The two-ranked leaves of Potentilla glaucophylla suggest a possible connection to P. crantzii in sect. Aureae. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. Potentilla hickmanii is known from only two extant populations in Monterey and San Mateo counties. The species is a federally listed endangered species (U.S. Fish and Wildlife Service, ecos.fws.gov/docs/recovery_plan/050617a.pdf). Plants from Sonoma County formerly included in P. hickmanii now constitute P. uliginosa. Although some morphologic overlap occurs with P. millefolia, P. hickmanii generally differs in having less dissected leaflets occupying less of the leaf axis, as well as somewhat larger flowers bearing fewer carpels. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 9, p. 152. | FNA vol. 9, p. 174. | ||||
Parent taxa | Rosaceae > subfam. Rosoideae > tribe Potentilleae > Potentilla > sect. Graciles | Rosaceae > subfam. Rosoideae > tribe Potentilleae > Potentilla > sect. Multijugae | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | P. diversifolia var. glaucophylla | |||||
Name authority | Lehmann: Index Seminum (Hamburg) 1836: 7. (1836) | Eastwood: Bull. Torrey Bot. Club 29: 77. (1902) | ||||
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