Potentilla concinna |
Potentilla uliginosa |
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early cinquefoil, elegant cinquefoil, red cinquefoil |
Cunningham Marsh cinquefoil |
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Habit | Plants rosetted to tufted; taproots not fleshy-thickened. | |||||||||
Stems | 0.2–1.6 dm, lengths 1/2–3(–4) times basal leaves. |
probably prostrate to decumbent, or ± ascending in supporting vegetation, 2.5–5.5 dm, lengths 1–2 times basal leaves. |
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Basal leaves | usually palmate to subpalmate, sometimes subpinnate, 1(–2) pair of leaflets separate from terminal leaflets, distal leaflets distinct, 2–10(–15) cm; petiole 1–7(–10) cm, straight hairs ± abundant, ± appressed, 1–3 mm, stiff to weak, cottony hairs present, sometimes absent; leaflets 5(–7), on tip or to distal 1/4 of leaf axis, separate to ± overlapping, proximal pair separated from others by 0–5(–10) mm of leaf axis, central leaflets narrowly oblanceolate to obovate, 1–3(–7) × 0.4–1.2 cm, petiolules 0–2(–4) mm, distal 1/4 to whole margin incised 1/4–3/4+ to midvein, teeth (1–)2–5(–10) per side, separate, 1–6 mm, surfaces strongly to ± dissimilar, abaxial grayish to white, straight hairs ± abundant, ± appressed, 0.5–2 mm, weak to stiff (especially on veins), cottony hairs ± dense, rarely sparse, glands sparse or obscured, adaxial green to grayish, straight hairs common to abundant, appressed, 0.5–2 mm, mostly stiff, sometimes weak or mixed, cottony hairs absent or sparse, rarely common, glands ± sparse. |
pinnate with distal leaflets ± confluent, 15–32 × 2–4 cm; petiole 5–10(–15) cm, straight hairs absent or sparse to common, appressed, 0.5–1.5 mm, stiff, cottony hairs absent, glands absent or sparse; primary lateral leaflets 6–10(–12) per side (irregularly paired), on distal ± 1/2 of leaf axis, loosely overlapping to proximally separate, largest ones cuneate to flabellate, 1.2–2.2 × 1–3 cm, ± whole margin unevenly incised 3/4 to nearly to midvein, ultimate segments (3–)5–10(–15), linear to narrowly oblanceolate, 5–17 × 1–2.5 mm, apical tufts less than 0.5 mm, surfaces green, not glaucous, straight hairs sparse (often glabrate adaxially), appressed, 0.5–1(–1.5) mm, stiff, cottony hairs absent, glands absent or sparse. |
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Cauline leaves | 2. |
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Inflorescences | (1–)2–12-flowered. |
6–10-flowered, very openly cymose. |
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Pedicels | 1–2(–3) cm. |
1–6 cm, ± recurved in fruit. |
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Flowers | epicalyx bractlets lanceolate to narrowly ovate to linear-elliptic, sometimes doubled, (2–)2.5–5 × 1–1.5 mm; hypanthium 3–6 mm diam.; sepals 3.5–6 mm, apex acute; petals (2.5–)4–9 × (2–)3–7 mm; filaments 1–3 mm, anthers 0.5–1 mm; carpels (7–)10–30, styles (1–)1.5–2 mm. |
epicalyx bractlets narrowly elliptic to oblong-ovate, 4–6 × 1–2.5 mm; hypanthium 5–6 mm diam.; sepals 4–7 mm, apex acute; petals 6–10 × 5–8 mm; filaments (1.5–)2–3 mm, anthers 0.7–1.2 mm; carpels ca. 10, styles 2.5–3.5 mm. |
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Achenes | 1.5–2.5 mm, smooth to lightly rugose. |
2–2.6 mm, smooth, ± carunculate. |
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2n | = 70. |
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Potentilla concinna |
Potentilla uliginosa |
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Phenology | Flowering spring–summer. | |||||||||
Habitat | Permanent oligotrophic wetlands | |||||||||
Elevation | 30–40 m (100–100 ft) | |||||||||
Distribution |
AZ; CA; CO; MT; ND; NM; NV; OH; SD; UT; WY; AB; MB; SK
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CA |
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Discussion | Varieties 3 (3 in the flora). Potentilla concinna is a relatively widespread, low-growing, early-blooming species with palmate to subpalmate, abaxially cottony leaves. Flowers often appear before leaves are fully expanded. The three varieties treated here have some level of biogeographic identity and are, therefore, given provisional taxonomic recognition. Some treatments have also included P. bicrenata, P. johnstonii, and P. macounii as varieties of P. concinna. Two Mexican species, P. leonina Standley and P. oblanceolata Rydberg, have also been treated as varieties of P. concinna by J. Soják (2006); they are retained as distinct species here, pending further research on Mexican Potentilla. Two other erstwhile varieties of P. concinna are now placed in separate sections: var. modesta (Rydberg) S. L. Welsh & B. C. Johnston (misapplied) in sect. Rubricaules as P. modesta, and var. rubripes (Rydberg) C. L. Hitchcock in sect. Subjugae, intermediate between P. saximontana and P. subjuga. Potentilla concinnaeformis Rydberg, considered to be a close relative of P. concinna by I. W. Clokey (1939), is more likely a hybrid between P. glaucophylla (sect. Graciles) and P. hippiana (sect. Leucophyllae). Hybrids also occur with P. multisecta, and probably other species as well. Potentilla concinna was first described in 1818 as P. humifusa Nuttall, a later homonym for P. humifusa Willdenow. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. Potentilla uliginosa is known only from a handful of historical collections from Cunningham Marsh, one of three wetlands in southern Sonoma County that harbor a suite of disjunct and regionally rare species (B. C. Johnston and B. Ertter 2010). Although previously included in P. hickmanii, P. uliginosa differs in having woodier caudices, longer stems and leaves, and more deeply incised leaflets occupying less of the leaf axis. Potentilla uliginosa is presumed extinct; however, a morphologically comparable collection of the P. millefolia complex from Josephine County, Oregon (Deer Creek, near Selma, 14 June 1929, Applegate 5735, UC), opens the possibility of additional undiscovered populations in the mountains of northwestern California. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 9, p. 178. | FNA vol. 9, p. 175. | ||||||||
Parent taxa | Rosaceae > subfam. Rosoideae > tribe Potentilleae > Potentilla > sect. Concinnae | Rosaceae > subfam. Rosoideae > tribe Potentilleae > Potentilla > sect. Multijugae | ||||||||
Sibling taxa | ||||||||||
Subordinate taxa | ||||||||||
Name authority | Richardson: in J. Franklin, Narr. Journey Polar Sea, 739. (1823) | B. C. Johnston & Ertter: J. Bot. Res. Inst. Texas 4: 14, fig. 1. (2010) | ||||||||
Web links |