Potentilla concinna
Potentilla nana
early cinquefoil, elegant cinquefoil, red cinquefoil
arctic cinquefoil, dwarf cinquefoil
0.2–1.6 dm, lengths 1/2–3(–4) times basal leaves.
ascending to erect, 0.1–0.5(–0.7) dm, lengths 1–2 times basal leaves.
usually palmate to subpalmate, sometimes subpinnate, 1(–2) pair of leaflets separate from terminal leaflets, distal leaflets distinct, 2–10(–15) cm;
petiole 1–7(–10) cm, straight hairs ± abundant, ± appressed, 1–3 mm, stiff to weak, cottony hairs present, sometimes absent;
leaflets 5(–7), on tip or to distal 1/4 of leaf axis, separate to ± overlapping, proximal pair separated from others by 0–5(–10) mm of leaf axis, central leaflets narrowly oblanceolate to obovate, 1–3(–7) × 0.4–1.2 cm, petiolules 0–2(–4) mm, distal 1/4 to whole margin incised 1/4–3/4+ to midvein, teeth (1–)2–5(–10) per side, separate, 1–6 mm, surfaces strongly to ± dissimilar, abaxial grayish to white, straight hairs ± abundant, ± appressed, 0.5–2 mm, weak to stiff (especially on veins), cottony hairs ± dense, rarely sparse, glands sparse or obscured, adaxial green to grayish, straight hairs common to abundant, appressed, 0.5–2 mm, mostly stiff, sometimes weak or mixed, cottony hairs absent or sparse, rarely common, glands ± sparse.
not in ranks, ternate, 1–5 cm;
stipules: apex acute;
petiole 0.5–3.5 cm, long hairs sparse to abundant, ascending to spreading, 0.5–2 mm, ± soft, glands sparse to common;
leaflets 3, central obovate, 0.5–2 × 0.5–1 cm, petiolule 0–1 mm, margins revolute, not lobed, distal 1/2–2/3 evenly incised ± 1/2 to midvein, teeth 3–4(–6) per side, surfaces ± similar, green (paler abaxially), hairs sparse to abundant, 0.8–1.2 mm, glands sparse to common.
(1–)2–12-flowered.
1(–2)-flowered.
1–2(–3) cm.
straight, 0.2–1.5 cm in flower, to 5 cm in fruit.
epicalyx bractlets lanceolate to narrowly ovate to linear-elliptic, sometimes doubled, (2–)2.5–5 × 1–1.5 mm;
hypanthium 3–6 mm diam.;
sepals 3.5–6 mm, apex acute;
petals (2.5–)4–9 × (2–)3–7 mm;
filaments 1–3 mm, anthers 0.5–1 mm;
carpels (7–)10–30, styles (1–)1.5–2 mm.
epicalyx bractlets oblong or ovate, 2.5–5 × 1.5–3.5 mm, margins revolute;
hypanthium 3–3.5 mm diam.;
sepals 2.5–5 mm, apex ± acute;
petals pale yellow, 4–8 × 4–6 mm;
filaments 1.2–2 mm, anthers 0.3–0.4 mm;
carpels 40–50, styles ± columnar, not or scarcely papillate-swollen proximally, 0.8–1.2 mm.
1.5–2.5 mm, smooth to lightly rugose.
1.4–1.6 mm.
= 70.
Potentilla concinna
Potentilla nana
Varieties 3 (3 in the flora).
Potentilla concinna is a relatively widespread, low-growing, early-blooming species with palmate to subpalmate, abaxially cottony leaves. Flowers often appear before leaves are fully expanded. The three varieties treated here have some level of biogeographic identity and are, therefore, given provisional taxonomic recognition. Some treatments have also included P. bicrenata, P. johnstonii, and P. macounii as varieties of P. concinna. Two Mexican species, P. leonina Standley and P. oblanceolata Rydberg, have also been treated as varieties of P. concinna by J. Soják (2006); they are retained as distinct species here, pending further research on Mexican Potentilla. Two other erstwhile varieties of P. concinna are now placed in separate sections: var. modesta (Rydberg) S. L. Welsh & B. C. Johnston (misapplied) in sect. Rubricaules as P. modesta, and var. rubripes (Rydberg) C. L. Hitchcock in sect. Subjugae, intermediate between P. saximontana and P. subjuga. Potentilla concinnaeformis Rydberg, considered to be a close relative of P. concinna by I. W. Clokey (1939), is more likely a hybrid between P. glaucophylla (sect. Graciles) and P. hippiana (sect. Leucophyllae). Hybrids also occur with P. multisecta, and probably other species as well.
Potentilla concinna was first described in 1818 as P. humifusa Nuttall, a later homonym for P. humifusa Willdenow.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Potentilla nana has most often been considered a race of P. hyparctica or as the priority name for P. hyparctica. As now defined, P. nana occurs in an arc across southern Alaska, from the Aleutian and Pribilof islands to the Alaskan panhandle and adjacent British Columbia. T. Wolf (1908) treated it as a dwarf form of P. fragiformis; J. Soják (1996) interpreted P. nana as a hybrid species originating from cross(es) between P. fragiformis and P. hyparctica. Although the hybrid hypothesis has morphologic support, P. nana has a unique combination of morphologic features and has a distinct range from both P. fragiformis and P. hyparctica. The species are also ecologically segregated: P. nana is a coastal heath and gravel slope species, like P. hyparctica; P. fragiformis is strictly maritime coastal.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
1. Distal (1/2–)3/4 to full length of central leaflets incised 1/2–3/4+ to midvein, teeth 2–6 mm; n Great Plains to c Utah. | var. divisa |
1. Distal 1/4–3/4 of central leaflets incised 1/4–1/2 to midvein, teeth 1–3(–5) mm; mostly Rocky Mountains and west | → 2 |
2. Stem lengths 1/2–1.5(–2) times basal leaves; leaves palmate, proximal leaflet pair not separated from others. | var. concinna |
2. Stem lengths 2–3(–4) times basal leaves; leaves usually palmate to subpalmate, sometimes subpinnate, proximal leaflet pair often separated from others by 1–5(–10) mm of leaf axis. | var. proxima |
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