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early cinquefoil, elegant cinquefoil, red cinquefoil

Jepson's cinquefoil, Jepson's potentilla

Stems

0.2–1.6 dm, lengths 1/2–3(–4) times basal leaves.

decumbent to nearly erect, (0.2–)0.5–2.6 dm.

Basal leaves

usually palmate to subpalmate, sometimes subpinnate, 1(–2) pair of leaflets separate from terminal leaflets, distal leaflets distinct, 2–10(–15) cm;

petiole 1–7(–10) cm, straight hairs ± abundant, ± appressed, 1–3 mm, stiff to weak, cottony hairs present, sometimes absent;

leaflets 5(–7), on tip or to distal 1/4 of leaf axis, separate to ± overlapping, proximal pair separated from others by 0–5(–10) mm of leaf axis, central leaflets narrowly oblanceolate to obovate, 1–3(–7) × 0.4–1.2 cm, petiolules 0–2(–4) mm, distal 1/4 to whole margin incised 1/4–3/4+ to midvein, teeth (1–)2–5(–10) per side, separate, 1–6 mm, surfaces strongly to ± dissimilar, abaxial grayish to white, straight hairs ± abundant, ± appressed, 0.5–2 mm, weak to stiff (especially on veins), cottony hairs ± dense, rarely sparse, glands sparse or obscured, adaxial green to grayish, straight hairs common to abundant, appressed, 0.5–2 mm, mostly stiff, sometimes weak or mixed, cottony hairs absent or sparse, rarely common, glands ± sparse.

subpinnate to subpalmate, 1.5–10(–17) cm;

petiole 0.5–4(–8) cm, long hairs ± abundant, spreading to loosely appressed, 0.5–2 mm, weak to stiff, short hairs absent or sparse, cottony or crisped hairs absent, glands sparse to abundant;

leaflets (1–)2–3 per side, on distal 1/6–1/3(–1/2) of leaf axis, separate to slightly overlapping, terminal ones oblanceolate-elliptic to narrowly obovate, (0.5–)1–2.5(–3.5) × 0.4–1(–1.3) cm, margins strongly revolute, incised 3/4+ to midvein, undivided medial blade 1.5–3(–5) mm wide, teeth (2–)3–5(–6) per side, ± linear-elliptic to rarely ovate or oblanceolate, surfaces ± dissimilar, abaxial ± grayish, rarely white, long hairs ± abundant especially on veins, 1 mm, weak, short hairs absent or sparse, cottony hairs absent, crisped hairs sparse to abundant, glands ± abundant, sometimes obscured, adaxial green to gray-green, long hairs sparse to abundant, loosely appressed, 0.5–1(–2) mm, short hairs absent or sparse, cottony and crisped hairs absent, glands ± abundant.

Cauline leaves

1–2.

Inflorescences

(1–)2–12-flowered.

(1–)3–12(–20)-flowered, congested, scarcely opening in fruit.

Pedicels

1–2(–3) cm.

0.2–0.5 cm (proximal to 1.2 cm).

Flowers

epicalyx bractlets lanceolate to narrowly ovate to linear-elliptic, sometimes doubled, (2–)2.5–5 × 1–1.5 mm;

hypanthium 3–6 mm diam.;

sepals 3.5–6 mm, apex acute;

petals (2.5–)4–9 × (2–)3–7 mm;

filaments 1–3 mm, anthers 0.5–1 mm;

carpels (7–)10–30, styles (1–)1.5–2 mm.

epicalyx bractlets elliptic to linear, 1.5–3 × (0.3–)0.5–1.2 mm, lengths 1/2–3/4 times sepals, margins flat;

hypanthium 2.5–5 mm diam.;

sepals 2–5 mm, apex usually ± acute to obtuse, rarely acuminate, abaxial surfaces: venation faint, glands abundant, evident;

petals pale yellow, 2–5 × 2–4 mm, lengths ± equal to sepals;

filaments 0.5–2 mm, anthers 0.3–0.7 mm;

carpels 20–60, styles papillate-swollen in proximal 1/2–3/4+, 0.8–1.2 mm.

Achenes

1.5–2.5 mm, smooth to lightly rugose.

1 mm, smooth to faintly rugose.

2n

= 70.

Potentilla concinna

Potentilla jepsonii

Phenology Flowering summer.
Habitat Seasonally moist sites in alpine meadows, ridges, fellfields, rock ledges, talus slopes
Elevation 2500–3700 m (8200–12100 ft)
Distribution
from FNA
AZ; CA; CO; MT; ND; NM; NV; OH; SD; UT; WY; AB; MB; SK
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; CA; CO; ID; MT; NV; OR; UT; WA; AB
[BONAP county map]
Discussion

Varieties 3 (3 in the flora).

Potentilla concinna is a relatively widespread, low-growing, early-blooming species with palmate to subpalmate, abaxially cottony leaves. Flowers often appear before leaves are fully expanded. The three varieties treated here have some level of biogeographic identity and are, therefore, given provisional taxonomic recognition. Some treatments have also included P. bicrenata, P. johnstonii, and P. macounii as varieties of P. concinna. Two Mexican species, P. leonina Standley and P. oblanceolata Rydberg, have also been treated as varieties of P. concinna by J. Soják (2006); they are retained as distinct species here, pending further research on Mexican Potentilla. Two other erstwhile varieties of P. concinna are now placed in separate sections: var. modesta (Rydberg) S. L. Welsh & B. C. Johnston (misapplied) in sect. Rubricaules as P. modesta, and var. rubripes (Rydberg) C. L. Hitchcock in sect. Subjugae, intermediate between P. saximontana and P. subjuga. Potentilla concinnaeformis Rydberg, considered to be a close relative of P. concinna by I. W. Clokey (1939), is more likely a hybrid between P. glaucophylla (sect. Graciles) and P. hippiana (sect. Leucophyllae). Hybrids also occur with P. multisecta, and probably other species as well.

Potentilla concinna was first described in 1818 as P. humifusa Nuttall, a later homonym for P. humifusa Willdenow.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Potentilla jepsonii (a replacement name for P. pensylvanica var. ovium) is widespread at higher elevations in the Intermountain Region, extending into the Sierra Nevada in California and the Rocky Mountains in Idaho and Wyoming. Disjunct occurrences are known on the north rim of the Grand Canyon and San Francisco Peaks, Arizona, and Steens Mountain, Oregon. Collections have most commonly been included in P. pensylvanica (for example, B. Ertter 1993; N. H. Holmgren 1997b); they differ in leaf division, vestiture, and epicalyx bractlets. J. Soják considered var. ovium to be a variety of P. litoralis, which has comparable leaves and vestiture; P. jepsonii lacks the enlarged, revolute epicalyx bractlets that distinguish P. litoralis and is for the most part well separated geographically and elevationally. The exception is the montane phase of P. litoralis, which, at the current stage of knowledge, is somewhat arbitrarily delimited from P. jepsonii in the northern Rocky Mountains; collections from Waterton Lakes National Park in Alberta and the Okanogan Mountains in Washington are tentatively placed here in P. jepsonii.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Distal (1/2–)3/4 to full length of central leaflets incised 1/2–3/4+ to midvein, teeth 2–6 mm; n Great Plains to c Utah.
var. divisa
1. Distal 1/4–3/4 of central leaflets incised 1/4–1/2 to midvein, teeth 1–3(–5) mm; mostly Rocky Mountains and west
→ 2
2. Stem lengths 1/2–1.5(–2) times basal leaves; leaves palmate, proximal leaflet pair not separated from others.
var. concinna
2. Stem lengths 2–3(–4) times basal leaves; leaves usually palmate to subpalmate, sometimes subpinnate, proximal leaflet pair often separated from others by 1–5(–10) mm of leaf axis.
var. proxima
Source FNA vol. 9, p. 178. FNA vol. 9, p. 216.
Parent taxa Rosaceae > subfam. Rosoideae > tribe Potentilleae > Potentilla > sect. Concinnae Rosaceae > subfam. Rosoideae > tribe Potentilleae > Potentilla > sect. Pensylvanicae
Sibling taxa
P. albiflora, P. ambigens, P. anachoretica, P. angelliae, P. anglica, P. anserina, P. arenosa, P. argentea, P. arizonica, P. basaltica, P. bicrenata, P. biennis, P. biflora, P. bimundorum, P. bipinnatifida, P. brevifolia, P. breweri, P. bruceae, P. brunnescens, P. canadensis, P. cottamii, P. crantzii, P. crebridens, P. crinita, P. cristae, P. demotica, P. drummondii, P. effusa, P. elegans, P. erecta, P. flabellifolia, P. fragiformis, P. furcata, P. glaucophylla, P. gracilis, P. grayi, P. hickmanii, P. hippiana, P. holmgrenii, P. hookeriana, P. hyparctica, P. inclinata, P. intermedia, P. jepsonii, P. johnstonii, P. lasiodonta, P. litoralis, P. macounii, P. millefolia, P. modesta, P. morefieldii, P. multijuga, P. multisecta, P. nana, P. newberryi, P. nivea, P. norvegica, P. ovina, P. paucijuga, P. pedersenii, P. pensylvanica, P. plattensis, P. pseudosericea, P. pulchella, P. pulcherrima, P. recta, P. reptans, P. rhyolitica, P. rimicola, P. rivalis, P. robbinsiana, P. rubella, P. rubricaulis, P. sanguinea, P. saximontana, P. sierrae-blancae, P. simplex, P. sterilis, P. stipularis, P. subgorodkovii, P. subjuga, P. subvahliana, P. subviscosa, P. supina, P. thurberi, P. thuringiaca, P. tikhomirovii, P. townsendii, P. uliginosa, P. uschakovii, P. vahliana, P. verna, P. versicolor, P. villosa, P. villosula, P. vulcanicola, P. wheeleri
P. albiflora, P. ambigens, P. anachoretica, P. angelliae, P. anglica, P. anserina, P. arenosa, P. argentea, P. arizonica, P. basaltica, P. bicrenata, P. biennis, P. biflora, P. bimundorum, P. bipinnatifida, P. brevifolia, P. breweri, P. bruceae, P. brunnescens, P. canadensis, P. concinna, P. cottamii, P. crantzii, P. crebridens, P. crinita, P. cristae, P. demotica, P. drummondii, P. effusa, P. elegans, P. erecta, P. flabellifolia, P. fragiformis, P. furcata, P. glaucophylla, P. gracilis, P. grayi, P. hickmanii, P. hippiana, P. holmgrenii, P. hookeriana, P. hyparctica, P. inclinata, P. intermedia, P. johnstonii, P. lasiodonta, P. litoralis, P. macounii, P. millefolia, P. modesta, P. morefieldii, P. multijuga, P. multisecta, P. nana, P. newberryi, P. nivea, P. norvegica, P. ovina, P. paucijuga, P. pedersenii, P. pensylvanica, P. plattensis, P. pseudosericea, P. pulchella, P. pulcherrima, P. recta, P. reptans, P. rhyolitica, P. rimicola, P. rivalis, P. robbinsiana, P. rubella, P. rubricaulis, P. sanguinea, P. saximontana, P. sierrae-blancae, P. simplex, P. sterilis, P. stipularis, P. subgorodkovii, P. subjuga, P. subvahliana, P. subviscosa, P. supina, P. thurberi, P. thuringiaca, P. tikhomirovii, P. townsendii, P. uliginosa, P. uschakovii, P. vahliana, P. verna, P. versicolor, P. villosa, P. villosula, P. vulcanicola, P. wheeleri
Subordinate taxa
P. concinna var. concinna, P. concinna var. divisa, P. concinna var. proxima
Synonyms P. pensylvanica var. ovium, P. litoralis var. ovium
Name authority Richardson: in J. Franklin, Narr. Journey Polar Sea, 739. (1823) Ertter: J. Bot. Res. Inst. Texas 2: 202. (2008)
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