Potentilla concinna |
Potentilla hickmanii |
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early cinquefoil, elegant cinquefoil, red cinquefoil |
Hickman's cinquefoil |
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Habit | Plants ± rosetted; taproots fleshy-thickened. | |||||||||
Stems | 0.2–1.6 dm, lengths 1/2–3(–4) times basal leaves. |
usually prostrate to decumbent, sometimes ± ascending in supporting vegetation, 0.5–2.5 dm, lengths 1–2 times basal leaves. |
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Basal leaves | usually palmate to subpalmate, sometimes subpinnate, 1(–2) pair of leaflets separate from terminal leaflets, distal leaflets distinct, 2–10(–15) cm; petiole 1–7(–10) cm, straight hairs ± abundant, ± appressed, 1–3 mm, stiff to weak, cottony hairs present, sometimes absent; leaflets 5(–7), on tip or to distal 1/4 of leaf axis, separate to ± overlapping, proximal pair separated from others by 0–5(–10) mm of leaf axis, central leaflets narrowly oblanceolate to obovate, 1–3(–7) × 0.4–1.2 cm, petiolules 0–2(–4) mm, distal 1/4 to whole margin incised 1/4–3/4+ to midvein, teeth (1–)2–5(–10) per side, separate, 1–6 mm, surfaces strongly to ± dissimilar, abaxial grayish to white, straight hairs ± abundant, ± appressed, 0.5–2 mm, weak to stiff (especially on veins), cottony hairs ± dense, rarely sparse, glands sparse or obscured, adaxial green to grayish, straight hairs common to abundant, appressed, 0.5–2 mm, mostly stiff, sometimes weak or mixed, cottony hairs absent or sparse, rarely common, glands ± sparse. |
pinnate with distal leaflets ± confluent, 3–17 × 1–3.5 cm; petiole 1–5 cm, straight hairs sparse to common, appressed, 0.5–1.5(–2) mm, stiff, cottony hairs absent, glands absent or sparse; primary lateral leaflets 3–6 per side, on distal 1/2–2/3 of leaf axis, ± overlapping to nearly separate, largest ones cuneate to flabellate, 0.5–2 × 0.5–1.5 cm, distal 1/2 to whole margin ± palmately incised 1/2–2/3(+) to midvein, ultimate teeth or segments 2–5, narrowly elliptic to oblanceolate, 2–10(–15) × (1–)1.5–3 mm, apical tufts 0.5 mm, surfaces green, not glaucous, straight hairs sparse to common (often sparser to glabrate adaxially), appressed, 1–1.5 mm, stiff, cottony hairs absent, glands absent or inconspicuous. |
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Cauline leaves | 2. |
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Inflorescences | (1–)2–12-flowered. |
2–5-flowered, very openly cymose, sometimes racemiform. |
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Pedicels | 1–2(–3) cm. |
1–2(–4) cm, ± recurved in fruit. |
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Flowers | epicalyx bractlets lanceolate to narrowly ovate to linear-elliptic, sometimes doubled, (2–)2.5–5 × 1–1.5 mm; hypanthium 3–6 mm diam.; sepals 3.5–6 mm, apex acute; petals (2.5–)4–9 × (2–)3–7 mm; filaments 1–3 mm, anthers 0.5–1 mm; carpels (7–)10–30, styles (1–)1.5–2 mm. |
epicalyx bractlets ± elliptic, 3.5–6 × (1–)1.5–2.5 mm; hypanthium 3–5 mm diam.; sepals 3.5–7(–9) mm, apex acute; petals 6–12 × 5–9 mm; filaments (1.5–)2–3(–4) mm, anthers 0.7–1.2 mm; carpels (2–)5–15, styles 2.5–3.5 mm. |
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Achenes | 1.5–2.5 mm, smooth to lightly rugose. |
2 mm, smooth, ± carunculate. |
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2n | = 70. |
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Potentilla concinna |
Potentilla hickmanii |
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Phenology | Flowering spring–early summer. | |||||||||
Habitat | Vernally saturated coastal meadows, openings in Monterey Pine forests | |||||||||
Elevation | 30–100 m (100–300 ft) | |||||||||
Distribution |
AZ; CA; CO; MT; ND; NM; NV; OH; SD; UT; WY; AB; MB; SK
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CA |
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Discussion | Varieties 3 (3 in the flora). Potentilla concinna is a relatively widespread, low-growing, early-blooming species with palmate to subpalmate, abaxially cottony leaves. Flowers often appear before leaves are fully expanded. The three varieties treated here have some level of biogeographic identity and are, therefore, given provisional taxonomic recognition. Some treatments have also included P. bicrenata, P. johnstonii, and P. macounii as varieties of P. concinna. Two Mexican species, P. leonina Standley and P. oblanceolata Rydberg, have also been treated as varieties of P. concinna by J. Soják (2006); they are retained as distinct species here, pending further research on Mexican Potentilla. Two other erstwhile varieties of P. concinna are now placed in separate sections: var. modesta (Rydberg) S. L. Welsh & B. C. Johnston (misapplied) in sect. Rubricaules as P. modesta, and var. rubripes (Rydberg) C. L. Hitchcock in sect. Subjugae, intermediate between P. saximontana and P. subjuga. Potentilla concinnaeformis Rydberg, considered to be a close relative of P. concinna by I. W. Clokey (1939), is more likely a hybrid between P. glaucophylla (sect. Graciles) and P. hippiana (sect. Leucophyllae). Hybrids also occur with P. multisecta, and probably other species as well. Potentilla concinna was first described in 1818 as P. humifusa Nuttall, a later homonym for P. humifusa Willdenow. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. Potentilla hickmanii is known from only two extant populations in Monterey and San Mateo counties. The species is a federally listed endangered species (U.S. Fish and Wildlife Service, ecos.fws.gov/docs/recovery_plan/050617a.pdf). Plants from Sonoma County formerly included in P. hickmanii now constitute P. uliginosa. Although some morphologic overlap occurs with P. millefolia, P. hickmanii generally differs in having less dissected leaflets occupying less of the leaf axis, as well as somewhat larger flowers bearing fewer carpels. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 9, p. 178. | FNA vol. 9, p. 174. | ||||||||
Parent taxa | Rosaceae > subfam. Rosoideae > tribe Potentilleae > Potentilla > sect. Concinnae | Rosaceae > subfam. Rosoideae > tribe Potentilleae > Potentilla > sect. Multijugae | ||||||||
Sibling taxa | ||||||||||
Subordinate taxa | ||||||||||
Name authority | Richardson: in J. Franklin, Narr. Journey Polar Sea, 739. (1823) | Eastwood: Bull. Torrey Bot. Club 29: 77. (1902) | ||||||||
Web links |