Potentilla concinna |
Potentilla basaltica |
|||||||||
---|---|---|---|---|---|---|---|---|---|---|
early cinquefoil, elegant cinquefoil, red cinquefoil |
Black Rock potentilla, Soldier Meadow or Black Rock cinquefoil, soldier meadows cinquefoil |
|||||||||
Habit | Plants ± rosetted; taproots fleshy-thickened. | |||||||||
Stems | 0.2–1.6 dm, lengths 1/2–3(–4) times basal leaves. |
± prostrate, sometimes ascending in supporting vegetation, 1.5–5 dm, lengths 2–3 times basal leaves. |
||||||||
Basal leaves | usually palmate to subpalmate, sometimes subpinnate, 1(–2) pair of leaflets separate from terminal leaflets, distal leaflets distinct, 2–10(–15) cm; petiole 1–7(–10) cm, straight hairs ± abundant, ± appressed, 1–3 mm, stiff to weak, cottony hairs present, sometimes absent; leaflets 5(–7), on tip or to distal 1/4 of leaf axis, separate to ± overlapping, proximal pair separated from others by 0–5(–10) mm of leaf axis, central leaflets narrowly oblanceolate to obovate, 1–3(–7) × 0.4–1.2 cm, petiolules 0–2(–4) mm, distal 1/4 to whole margin incised 1/4–3/4+ to midvein, teeth (1–)2–5(–10) per side, separate, 1–6 mm, surfaces strongly to ± dissimilar, abaxial grayish to white, straight hairs ± abundant, ± appressed, 0.5–2 mm, weak to stiff (especially on veins), cottony hairs ± dense, rarely sparse, glands sparse or obscured, adaxial green to grayish, straight hairs common to abundant, appressed, 0.5–2 mm, mostly stiff, sometimes weak or mixed, cottony hairs absent or sparse, rarely common, glands ± sparse. |
pinnate with distal leaflets ± confluent, 5–12(–18) × 1–1.5(–2) cm; petiole 0.5–1(–1.5) cm, straight hairs absent, cottony hairs absent, glands absent or sparse; primary lateral leaflets (5–)10–15(–20) per side, on nearly whole leaf axis, ± overlapping, largest ones (or leaflet lobes) elliptic, 0.4–1(–2) × 0.2–0.5(–1) cm, simple and entire or 2(–3)-fid to base (sometimes shallowly toothed as well), segments 1–2(–3), narrowly to broadly elliptic, 4–10(–20) × (1–)2–3 mm, apical tufts absent or less than 0.5 mm, surfaces green, glaucous, straight hairs absent or sparse to common, tightly appressed, 0.5 mm or less, stiff, cottony hairs absent, glands absent or sparse. |
||||||||
Cauline leaves | 2–4. |
|||||||||
Inflorescences | (1–)2–12-flowered. |
(5–)10–20-flowered, loosely cymose. |
||||||||
Pedicels | 1–2(–3) cm. |
0.8–3(–4) cm, straight to ± recurved in fruit. |
||||||||
Flowers | epicalyx bractlets lanceolate to narrowly ovate to linear-elliptic, sometimes doubled, (2–)2.5–5 × 1–1.5 mm; hypanthium 3–6 mm diam.; sepals 3.5–6 mm, apex acute; petals (2.5–)4–9 × (2–)3–7 mm; filaments 1–3 mm, anthers 0.5–1 mm; carpels (7–)10–30, styles (1–)1.5–2 mm. |
epicalyx bractlets lanceolate-elliptic to ovate-elliptic, 1.5–2.5(–3) × 1 mm; hypanthium 2–5 mm diam.; sepals (3–)4–5(–6) mm, apex acute to acuminate; petals (3.5–)4.5–6.5 × 2.5–4 mm; filaments 1.5–3 mm, anthers 0.6–0.8 mm; carpels 3–10, styles 2–2.5 mm. |
||||||||
Achenes | 1.5–2.5 mm, smooth to lightly rugose. |
1.8–2.2 mm, smooth to ± rugose, not carunculate. |
||||||||
2n | = 70. |
|||||||||
Potentilla concinna |
Potentilla basaltica |
|||||||||
Phenology | Flowering late spring–summer. | |||||||||
Habitat | Moist, subalkaline meadows in shrub steppe | |||||||||
Elevation | 1300–1600 m (4300–5200 ft) | |||||||||
Distribution |
AZ; CA; CO; MT; ND; NM; NV; OH; SD; UT; WY; AB; MB; SK
|
CA; NV |
||||||||
Discussion | Varieties 3 (3 in the flora). Potentilla concinna is a relatively widespread, low-growing, early-blooming species with palmate to subpalmate, abaxially cottony leaves. Flowers often appear before leaves are fully expanded. The three varieties treated here have some level of biogeographic identity and are, therefore, given provisional taxonomic recognition. Some treatments have also included P. bicrenata, P. johnstonii, and P. macounii as varieties of P. concinna. Two Mexican species, P. leonina Standley and P. oblanceolata Rydberg, have also been treated as varieties of P. concinna by J. Soják (2006); they are retained as distinct species here, pending further research on Mexican Potentilla. Two other erstwhile varieties of P. concinna are now placed in separate sections: var. modesta (Rydberg) S. L. Welsh & B. C. Johnston (misapplied) in sect. Rubricaules as P. modesta, and var. rubripes (Rydberg) C. L. Hitchcock in sect. Subjugae, intermediate between P. saximontana and P. subjuga. Potentilla concinnaeformis Rydberg, considered to be a close relative of P. concinna by I. W. Clokey (1939), is more likely a hybrid between P. glaucophylla (sect. Graciles) and P. hippiana (sect. Leucophyllae). Hybrids also occur with P. multisecta, and probably other species as well. Potentilla concinna was first described in 1818 as P. humifusa Nuttall, a later homonym for P. humifusa Willdenow. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. Potentilla basaltica is very distinctive in its glaucous, highly divided cylindric leaves with leaflets arranged in four ranks. The leaves strongly resemble those of some species of Ivesia, especially I. kingii S. Watson, but molecular analysis (B. Ertter et al. 1998) confirms a relationship with other members of sect. Multijugae. Potentilla basaltica is known from only two localities, one in Lassen County, California, and the other in Humboldt County, Nevada. Both populations are highly localized, and P. basaltica is a candidate for federal listing as well as of conservation concern in both states. The epithet basaltica was chosen as a reference to the Black Rock Desert, not habitat. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
||||||||
Key |
|
|||||||||
Source | FNA vol. 9, p. 178. | FNA vol. 9, p. 176. | ||||||||
Parent taxa | Rosaceae > subfam. Rosoideae > tribe Potentilleae > Potentilla > sect. Concinnae | Rosaceae > subfam. Rosoideae > tribe Potentilleae > Potentilla > sect. Multijugae | ||||||||
Sibling taxa | ||||||||||
Subordinate taxa | ||||||||||
Name authority | Richardson: in J. Franklin, Narr. Journey Polar Sea, 739. (1823) | Tiehm & Ertter: Brittonia 36: 228, fig. 1. (1984) | ||||||||
Web links |