Potentilla breweri
Potentilla crinita
Brewer's cinquefoil
bearded cinquefoil, Lemmon's cinquefoil
prostrate to ascending, (0.5–)1–3(–4.5) dm, lengths 2–4(–5) times basal leaves.
(0.5–)1.5–4.5 dm, lengths 2–4(–5) times basal leaves.
pinnate with distal leaflets ± confluent, (2–)4–12(–17) × 1–2.5(–5) cm;
petiole 1–3(–7) cm, straight hairs mostly absent, cottony hairs sparse to dense, glands absent or obscured to common;
primary lateral leaflets 3–6 per side, on distal 1/2–2/3(–3/4) of leaf axis, ± overlapping, largest ones cuneate-flabellate, 0.5–1.5(–2.5) × 0.4–2(–3) cm, distal 1/2 to whole margin unevenly incised 1/2 to completely to midvein (blade often medially split as well), ultimate teeth or segments 3–10, narrowly elliptic, 2–10(–15) × 1–3(–5) mm, apical tufts less than 1 mm, surfaces grayish green to white, not glaucous, straight hairs sparse to common (sparser adaxially), loosely appressed to ascending, 1–1.5 mm, soft, cottony hairs sparse to dense, glands sparse or obscured to common.
pinnate, 3–15(–20) cm;
petiole 1–10(–15) cm, long hairs dense, appressed, 1.5–2.5 mm, usually stiff, short and crisped hairs usually absent, cottony hairs absent, glands sparse, often obscured;
leaflets often conduplicate, lateral ones evenly paired, (3–)4–6(–7) per side on distal 1/3–2/3 of leaf axis, distal pairs ± decurrent, often confluent with terminal leaflet, larger leaflets narrowly cuneate or oblanceolate to obovate, 1–3(–4) × 0.2–0.8(–1) cm, distal 1/4–1/2(–2/3) or less of margin incised ± 1/4 or less to midvein, teeth (0–)1–5(–9) per side, 1–2 mm, surfaces ± similar to ± dissimilar, abaxial silvery to greenish, long hairs usually dense (at least on primary veins), 1–2 mm, stiff, short-crisped hairs absent or sparse, cottony hairs usually absent, glands sparse to common, often obscured, adaxial ± green, long hairs sparse to common, sometimes absent, short, crisped, and cottony hairs absent, glands sparse.
1–3.
1–3(–4).
2–15(–25)-flowered, openly cymose.
(5–)10–30-flowered.
(1–)1.5–2.5(–4) cm, straight in fruit.
0.5–2(–4) cm.
epicalyx bractlets lanceolate to ovate, 2–5 × 1–1.5 mm;
hypanthium (3–)4–5 mm diam.;
sepals (3–)4–7 mm, apex acute;
petals 5–9(–10) × 4–8(–10) mm;
filaments (1–)2–4 mm, anthers (0.5–)1 mm;
carpels 15–25, styles 2–3 mm.
epicalyx bractlets lanceolate, 1.5–4.5 × 0.5(–1) mm, 1/2–2/3 as long as sepals, abaxial vestiture similar to or ± sparser than sepals, not glabrescent, straight hairs common, crisped or cottony hairs usually absent;
hypanthium 2.5–5 mm diam.;
sepals (3–)4–7 mm, apex acute to long acuminate;
petals (3–)4.5–7.5(–8) × 4–6 mm;
filaments 1–3 mm, anthers 0.6–1.1 mm;
carpels 5–20, styles 1.6–2.6 mm.
1.8 mm, smooth, not carunculate.
1.4–1.7 mm, smooth or slightly rugose.
= 72–73, 99, 100, 102.
Potentilla breweri
Potentilla crinita
Of conservation concern.
Potentilla breweri differs from other members of sect. Multijugae in its often dense, cottony vestiture. Density of cottony hairs and leaflet dissection can differ significantly between first-formed and mid- to late-season leaves of P. breweri; in the description above, leaves are those predominant at anthesis.
Potentilla breweri is most common in the Sierra Nevada of California, with sporadic occurrences to the Cascade Range in southern Washington. Disjunct populations occur in the Ruby and Snake ranges in Nevada and on Steens Mountain in Oregon, where often introgressed with P. versicolor. Reports from Utah are based on collections of P. concinna var. proxima or possible hybrids between P. concinna var. proxima and P. ovina var. decurrens.
J. Clausen et al. (1940) concluded that Potentilla breweri (with P. versicolor as synonym) belonged with P. bruceae and P. drummondii as members of a cenospecies that probably also included unnamed species (possibly P. ovina var. decurrens) from the mountains of Colorado, Utah, and Wyoming. Chromosome number varied within populations, and high numbers of univalents were present.
Recognition of Potentilla breweri as a distinct species differs from recent treatments in which it was considered to be a variety or subspecies of P. drummondii, parallel to the treatment of P. bruceae. The present continental perspective of the genus, as well as additional fieldwork, has resulted in the return of all three to species rank (B. Ertter and D. Mansfield 2007). Placement of P. breweri in sect. Multijugae, separate from P. bruceae and P. drummondii in sect. Graciles, is based on the commonly prostrate habit, fully pinnate leaves, and deeply divided overlapping leaflets.
Although B. C. Johnston (1980) used Potentilla breweri var. viridis Jepson to accommodate greener-than-average sparsely cottony plants, including populations treated here as P. versicolor, the type is probably a sterile hybrid with P. wheeleri as one parent (B. Ertter 1992). The type of P. millefolia var. algida Jepson (included by Johnston in the synonymy of var. viridis) and comparable plants from north-central California combine features of P. breweri and P. versicolor but are more glandular than either. This variant, as well as exceptionally small plants from the Warner Mountains of California, may prove distinct.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Potentilla crinita occurs mainly in the upper foothills and mountains from southern Nevada to south-central Utah, northern Arizona, and northwestern New Mexico and is disjunct to southwestern Colorado (Archuleta County). It tends to grow on somewhat drier, rockier sites than co-occurring species of Potentilla. The often conduplicate leaflets, falcate in outline, bear relatively few, small teeth. Two varieties are sometimes recognized, based on leaflet and vestiture characters that do not reliably coincide.
Potentilla crinita can hybridize with P. hippiana where the two species overlap, in spite of ecological partitioning. N. H. Holmgren (1997b) noted the type of P. crinita may be such a hybrid. If correct, then P. lemmonii would be used for the species unless the name P. crinita were to be conserved with a conserved type.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
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