Potentilla bipinnatifida
Potentilla villosa
bipinnate cinquefoil, potentille bipinnatifide, tansy cinquefoil
hairy cinquefoil, northern cinquefoil, villous cinquefoil
stout, sometimes short-columnar, not sheathed with marcescent whole leaves.
ascending to erect, (1–)2–5 dm.
ascending, 0.5–2(–2.5) dm, lengths 1.5–3 times basal leaves.
subpinnate to subpalmate, (6–)10–25 cm;
petiole (2–)5–15 cm, long hairs dense, appressed, 1–2 mm, soft to ± stiff, short hairs absent, crisped hairs sparse, glands absent, sparse, or obscured;
leaflets 2–3 per side, on distal 1/6–1/3(–1/2) of leaf axis, separate to ± overlapping, terminal ones oblanceolate, (2–)3–6(–10) × 1–2(–3.5) cm, margins revolute, incised 3/4+ to midvein, undivided medial blade 1.5–6 mm wide, teeth 5–8 per side, ± linear, surfaces ± to strongly dissimilar, abaxial usually white, rarely grayish, long hairs abundant especially on veins, 1–2 mm, ± weak, short hairs absent or obscured, cottony (and crisped) hairs ± dense, glands absent or obscured, adaxial green to grayish, long hairs sparse to abundant, loosely appressed, 0.5–1.5 mm, short hairs absent or sparse, crisped and/or cottony hairs sparse to common, glands sparse to common.
2–12(–15) cm;
petiole 1–9(–12) cm, long hairs ± abundant to dense, spreading to ascending, 1–2.5(–3) mm, soft to weak, smooth, crisped hairs absent or sparse, cottony hairs absent, glands sparse to common, sometimes obscured;
leaflets usually ± overlapping, central broadly obovate to suborbiculate, (0.5–)1.5–2.5(–3) × (0.5–)1.5–2.6(–3.2) cm, sessile to subsessile, base cuneate to rounded, margins revolute, distal 1/2–2/3(–3/4) incised 1/4–1/2 to midvein, teeth 3–6(–7) per side, ± approximate to ± distant, surfaces ± dissimilar, abaxial yellowish or grayish white, long hairs 1–2 mm, cottony-crisped hairs ± dense, adaxial grayish green, long hairs abundant to dense, crisped hairs absent, sparse, or obscured.
2–4.
(0–)1–2.
(4–)10–50(–100)-flowered, congested or elongating in fruit.
(1–)2–7(–10)-flowered.
0.2–0.8 cm (proximal to 2 cm).
0.5–3(–4) cm in flower, to 4 cm in fruit.
epicalyx bractlets narrowly elliptic to lanceolate, 2.5–6 mm, lengths ± 2/3 times sepals, margins flat;
hypanthium 3–5 mm diam.;
sepals 3–6 mm, apex ± acute, abaxial surfaces: venation indistinct, glands absent, sparse, or obscured;
petals yellow, 3–5 × 3–4 mm, lengths ± equal to sepals;
filaments 0.5–2 mm, anthers 0.3–0.5 mm;
carpels 50–80, styles papillate-swollen in proximal 1/2–3/4+, 1–1.2 mm.
epicalyx bractlets ovate to oval-elliptic, 3–8 × 2–5 mm, 2/3 to as wide as sepals, margins strongly revolute, red glands absent;
hypanthium 5–7 mm diam.;
sepals 4–8 mm, apex ± acute;
petals (5–)7–15 × 7–16 mm, significantly longer than sepals;
filaments 1.8–2.1 mm, anthers 0.7–0.8 mm;
carpels 150–250, apical hairs usually absent, rarely present (cottony), styles narrowly conic to tapered, ± papillate-swollen on proximal 1/5–1/2, 0.9–1.1 mm.
1–1.2 mm, smooth to faintly rugose.
0.9–1.3 mm.
= 56.
= 14.
Potentilla bipinnatifida
Potentilla villosa
Potentilla bipinnatifida is similar to P. litoralis in habit and leaf dissection but has flat, silky epicalyx bractlets and sepals with no evident glands. Vestiture is generally silkier, and the silvery to bicolor leaves are white-cottony abaxially. The two species are sympatric in the plains of Canada, with some intergradation; P. bipinnatifida is also common south to Colorado, where it is found in intermontane meadows and sagebrush flats. Outlying populations occur in Blaine and Custer counties, Idaho, and Duchesne and Piute counties, Utah. Eastern collections from disturbed sites might be adventive.
Potentilla missourica Hornemann ex Lindley and P. normalis Besser ex Sprengel are older names for this species; both were rejected against a conserved P. bipinnatifida with designated lectotypes (see J. Soják 2008b).
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Potentilla villosa is the only documented diploid species of sect. Niveae in North America and, almost certainly, it is fully sexual. It is a characteristic species of coarse-grained beaches and coastal cliffs from southwestern British Columbia to western Alaska and the Russian Far East, and also occurs on scattered mountains in the Olympic Peninsula and Cascade Range of Washington and Oregon (providing the higher elevational extreme). These southern populations, which have been called var. parviflora, tend to be smaller, more delicate, less hairy plants in general, with fewer and smaller flowers. At least some plants in some southern populations also have cottony hairs on the carpels.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
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