Potentilla bipinnatifida
Potentilla versicolor
bipinnate cinquefoil, potentille bipinnatifide, tansy cinquefoil
Steens Mountain cinquefoil, varying cinquefoil
ascending to erect, (1–)2–5 dm.
prostrate to ascending, (0.7–)1.5–2.5(–4) dm, lengths 2–4(–5) times basal leaves.
subpinnate to subpalmate, (6–)10–25 cm;
petiole (2–)5–15 cm, long hairs dense, appressed, 1–2 mm, soft to ± stiff, short hairs absent, crisped hairs sparse, glands absent, sparse, or obscured;
leaflets 2–3 per side, on distal 1/6–1/3(–1/2) of leaf axis, separate to ± overlapping, terminal ones oblanceolate, (2–)3–6(–10) × 1–2(–3.5) cm, margins revolute, incised 3/4+ to midvein, undivided medial blade 1.5–6 mm wide, teeth 5–8 per side, ± linear, surfaces ± to strongly dissimilar, abaxial usually white, rarely grayish, long hairs abundant especially on veins, 1–2 mm, ± weak, short hairs absent or obscured, cottony (and crisped) hairs ± dense, glands absent or obscured, adaxial green to grayish, long hairs sparse to abundant, loosely appressed, 0.5–1.5 mm, short hairs absent or sparse, crisped and/or cottony hairs sparse to common, glands sparse to common.
pinnate with distal leaflets ± confluent, 4–12 × 1–2.5(–3.5) cm;
petiole 1–3 cm, straight hairs absent (on early-season petioles) or sparse to common, spreading-ascending to loosely appressed, 1–2 mm, soft, cottony hairs usually absent, glands sparse;
primary lateral leaflets 3–5 per side, on distal (1/3–)1/2–2/3 of leaf axis, overlapping, largest ones cuneate to flabellate, 0.5–1.5(–2) × 0.5–1.5 cm, distal 3/4 to whole margin unevenly incised 2/3 to completely to midvein (blade often medially split as well), ultimate teeth or segments 2–5(–8), ± oblanceolate, 3–11 × 1–3 mm, apical tufts to 1 mm, surfaces green to grayish green, not glaucous, straight hairs sparse to common (sparser adaxially), sometimes absent (except on margins), loosely appressed to ascending, 1–2 mm, soft, cottony hairs absent, glands sparse to common.
2–4.
1–2(–3).
(4–)10–50(–100)-flowered, congested or elongating in fruit.
(1–)3–10-flowered, usually openly cymose.
0.2–0.8 cm (proximal to 2 cm).
1–3(–5) cm, straight to slightly recurved in fruit.
epicalyx bractlets narrowly elliptic to lanceolate, 2.5–6 mm, lengths ± 2/3 times sepals, margins flat;
hypanthium 3–5 mm diam.;
sepals 3–6 mm, apex ± acute, abaxial surfaces: venation indistinct, glands absent, sparse, or obscured;
petals yellow, 3–5 × 3–4 mm, lengths ± equal to sepals;
filaments 0.5–2 mm, anthers 0.3–0.5 mm;
carpels 50–80, styles papillate-swollen in proximal 1/2–3/4+, 1–1.2 mm.
epicalyx bractlets lanceolate to elliptic, rarely ovate, 2–5 × 1–2 mm, sometimes apically toothed;
hypanthium 3–5 mm diam.;
sepals 4–7 mm, apex acute;
petals 4–7 × 3–5.5 mm;
filaments (1–)1.5–2.5 mm, anthers 0.5–0.8 mm;
carpels 10–25, styles 2 mm.
1–1.2 mm, smooth to faintly rugose.
1.5–1.8 mm, smooth to faintly rugose, not carunculate.
= 56.
Potentilla bipinnatifida
Potentilla versicolor
Potentilla bipinnatifida is similar to P. litoralis in habit and leaf dissection but has flat, silky epicalyx bractlets and sepals with no evident glands. Vestiture is generally silkier, and the silvery to bicolor leaves are white-cottony abaxially. The two species are sympatric in the plains of Canada, with some intergradation; P. bipinnatifida is also common south to Colorado, where it is found in intermontane meadows and sagebrush flats. Outlying populations occur in Blaine and Custer counties, Idaho, and Duchesne and Piute counties, Utah. Eastern collections from disturbed sites might be adventive.
Potentilla missourica Hornemann ex Lindley and P. normalis Besser ex Sprengel are older names for this species; both were rejected against a conserved P. bipinnatifida with designated lectotypes (see J. Soják 2008b).
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Potentilla versicolor is most common on Steens Mountain in Harney County, Oregon. Collections are known from other mountain ranges in eastern Oregon (some possibly representing distinct taxa) and from near Island Lake in the Ruby Mountains of northeastern Nevada. The species often grows and, apparently, intergrades with P. breweri; it is usually distinct in its lack of cottony hairs. Petals, filaments, and styles tend to be somewhat shorter in P. versicolor than in P. breweri. Collections of P. versicolor have most often been identified as P. breweri, P. millefolia, or P. ovina.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
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