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bipinnate cinquefoil, potentille bipinnatifide, tansy cinquefoil

Townsend's cinquefoil

Glands

absent or inconspicuous, uncolored.

Stems

ascending to erect, (1–)2–5 dm.

ascending to nearly erect, (1–)2–5(–7.5) dm.

Basal leaves

subpinnate to subpalmate, (6–)10–25 cm;

petiole (2–)5–15 cm, long hairs dense, appressed, 1–2 mm, soft to ± stiff, short hairs absent, crisped hairs sparse, glands absent, sparse, or obscured;

leaflets 2–3 per side, on distal 1/6–1/3(–1/2) of leaf axis, separate to ± overlapping, terminal ones oblanceolate, (2–)3–6(–10) × 1–2(–3.5) cm, margins revolute, incised 3/4+ to midvein, undivided medial blade 1.5–6 mm wide, teeth 5–8 per side, ± linear, surfaces ± to strongly dissimilar, abaxial usually white, rarely grayish, long hairs abundant especially on veins, 1–2 mm, ± weak, short hairs absent or obscured, cottony (and crisped) hairs ± dense, glands absent or obscured, adaxial green to grayish, long hairs sparse to abundant, loosely appressed, 0.5–1.5 mm, short hairs absent or sparse, crisped and/or cottony hairs sparse to common, glands sparse to common.

usually not in ranks, palmate, rarely subpalmate, (2–)5–20(–30) cm;

petiole (1–)2–15(–25) cm, long hairs usually absent or sparse and appressed, sometimes common and spreading, 1–1.5 mm, stiff, short, crisped, and cottony hairs absent, glands absent;

leaflets 5–7(–8), usually on tip, rarely on less than distal 1/10, of leaf axis, separate, largest ones narrowly oblanceolate-elliptic, (1.5–)2–5(–6) × 0.5–1.2(–1.5) cm, margins flat, distal (1/4–)1/2–3/4 evenly incised 1/4–1/3 to midvein, undivided medial blade 4–11 mm wide, teeth (2–)4–8(–9) per side, mostly broadly lanceolate, 1–2 mm, surfaces similar, green, not glaucous, long hairs usually absent or limited to veins and margins, short, crisped, and cottony hairs absent, glands absent.

Cauline leaves

2–4.

1–3.

Inflorescences

(4–)10–50(–100)-flowered, congested or elongating in fruit.

(5–)10–20(–25)-flowered.

Pedicels

0.2–0.8 cm (proximal to 2 cm).

0.7–3(–3.5) cm.

Flowers

epicalyx bractlets narrowly elliptic to lanceolate, 2.5–6 mm, lengths ± 2/3 times sepals, margins flat;

hypanthium 3–5 mm diam.;

sepals 3–6 mm, apex ± acute, abaxial surfaces: venation indistinct, glands absent, sparse, or obscured;

petals yellow, 3–5 × 3–4 mm, lengths ± equal to sepals;

filaments 0.5–2 mm, anthers 0.3–0.5 mm;

carpels 50–80, styles papillate-swollen in proximal 1/2–3/4+, 1–1.2 mm.

epicalyx bractlets lanceolate to elliptic, (2–)2.5–5(–6) × 0.5–1(–1.4) mm, hairs sparse, ± appressed, glands absent;

hypanthium 3.5–5 mm diam.;

sepals 4–8(–9) mm, apex acute to long acuminate;

petals 5–10 × 5–10(–12) mm;

filaments 1–3 mm, anthers (0.5–)0.6–1 mm;

carpels 15–30, styles filiform above papillate-swollen base, (1–)1.5–2 mm.

Achenes

1–1.2 mm, smooth to faintly rugose.

1.1–1.6 mm.

2n

= 56.

Potentilla bipinnatifida

Potentilla townsendii

Phenology Flowering summer. Flowering summer.
Habitat Open shortgrass prairie, alkaline bottoms, streamsides in sagebrush, disturbed sites Edges of meadows, dry gravelly flats and slopes, in montane to subalpine conifer woodlands, mixed grassland communities
Elevation 10–3400 m (0–11200 ft) (2300–)2700–3700 m ((7500–)8900–12100 ft)
Distribution
from FNA
AK; CO; ID; MI; MN; MT; ND; NE; NM; SD; UT; WY; AB; MB; NT; ON; QC; SK; YT
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; NM; Mexico (Chihuahua)
[BONAP county map]
Discussion

Potentilla bipinnatifida is similar to P. litoralis in habit and leaf dissection but has flat, silky epicalyx bractlets and sepals with no evident glands. Vestiture is generally silkier, and the silvery to bicolor leaves are white-cottony abaxially. The two species are sympatric in the plains of Canada, with some intergradation; P. bipinnatifida is also common south to Colorado, where it is found in intermontane meadows and sagebrush flats. Outlying populations occur in Blaine and Custer counties, Idaho, and Duchesne and Piute counties, Utah. Eastern collections from disturbed sites might be adventive.

Potentilla missourica Hornemann ex Lindley and P. normalis Besser ex Sprengel are older names for this species; both were rejected against a conserved P. bipinnatifida with designated lectotypes (see J. Soják 2008b).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Although previously applied only to collections from Chihuahua, Mexico, Potentilla townsendii is here used to accommodate most plants from Arizona and New Mexico previously placed in P. glaucophylla (as P. diversifolia Lehmann). Plants have narrow, shallowly toothed leaflets; stems, petioles, and leaflet veins that are often reddish; and larger anthers than P. glaucophylla. As now circumscribed, P. townsendii is known in the United States primarily from the White Mountains of east-central Arizona and the high mountains of northern New Mexico. Whether these plants are truly conspecific with those from Chihuahua, Mexico, remains to be confirmed. The species might also occur in Colorado and Utah, but the best disposition of these problematic populations is uncertain. Somewhat hairy plants with subpalmate leaves are interpreted as introgression from P. hippiana or other species.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 9, p. 217. FNA vol. 9, p. 153.
Parent taxa Rosaceae > subfam. Rosoideae > tribe Potentilleae > Potentilla > sect. Pensylvanicae Rosaceae > subfam. Rosoideae > tribe Potentilleae > Potentilla > sect. Graciles
Sibling taxa
P. albiflora, P. ambigens, P. anachoretica, P. angelliae, P. anglica, P. anserina, P. arenosa, P. argentea, P. arizonica, P. basaltica, P. bicrenata, P. biennis, P. biflora, P. bimundorum, P. brevifolia, P. breweri, P. bruceae, P. brunnescens, P. canadensis, P. concinna, P. cottamii, P. crantzii, P. crebridens, P. crinita, P. cristae, P. demotica, P. drummondii, P. effusa, P. elegans, P. erecta, P. flabellifolia, P. fragiformis, P. furcata, P. glaucophylla, P. gracilis, P. grayi, P. hickmanii, P. hippiana, P. holmgrenii, P. hookeriana, P. hyparctica, P. inclinata, P. intermedia, P. jepsonii, P. johnstonii, P. lasiodonta, P. litoralis, P. macounii, P. millefolia, P. modesta, P. morefieldii, P. multijuga, P. multisecta, P. nana, P. newberryi, P. nivea, P. norvegica, P. ovina, P. paucijuga, P. pedersenii, P. pensylvanica, P. plattensis, P. pseudosericea, P. pulchella, P. pulcherrima, P. recta, P. reptans, P. rhyolitica, P. rimicola, P. rivalis, P. robbinsiana, P. rubella, P. rubricaulis, P. sanguinea, P. saximontana, P. sierrae-blancae, P. simplex, P. sterilis, P. stipularis, P. subgorodkovii, P. subjuga, P. subvahliana, P. subviscosa, P. supina, P. thurberi, P. thuringiaca, P. tikhomirovii, P. townsendii, P. uliginosa, P. uschakovii, P. vahliana, P. verna, P. versicolor, P. villosa, P. villosula, P. vulcanicola, P. wheeleri
P. albiflora, P. ambigens, P. anachoretica, P. angelliae, P. anglica, P. anserina, P. arenosa, P. argentea, P. arizonica, P. basaltica, P. bicrenata, P. biennis, P. biflora, P. bimundorum, P. bipinnatifida, P. brevifolia, P. breweri, P. bruceae, P. brunnescens, P. canadensis, P. concinna, P. cottamii, P. crantzii, P. crebridens, P. crinita, P. cristae, P. demotica, P. drummondii, P. effusa, P. elegans, P. erecta, P. flabellifolia, P. fragiformis, P. furcata, P. glaucophylla, P. gracilis, P. grayi, P. hickmanii, P. hippiana, P. holmgrenii, P. hookeriana, P. hyparctica, P. inclinata, P. intermedia, P. jepsonii, P. johnstonii, P. lasiodonta, P. litoralis, P. macounii, P. millefolia, P. modesta, P. morefieldii, P. multijuga, P. multisecta, P. nana, P. newberryi, P. nivea, P. norvegica, P. ovina, P. paucijuga, P. pedersenii, P. pensylvanica, P. plattensis, P. pseudosericea, P. pulchella, P. pulcherrima, P. recta, P. reptans, P. rhyolitica, P. rimicola, P. rivalis, P. robbinsiana, P. rubella, P. rubricaulis, P. sanguinea, P. saximontana, P. sierrae-blancae, P. simplex, P. sterilis, P. stipularis, P. subgorodkovii, P. subjuga, P. subvahliana, P. subviscosa, P. supina, P. thurberi, P. thuringiaca, P. tikhomirovii, P. uliginosa, P. uschakovii, P. vahliana, P. verna, P. versicolor, P. villosa, P. villosula, P. vulcanicola, P. wheeleri
Synonyms P. pensylvanica var. bipinnatifida
Name authority Douglas: in W. J. Hooker, Fl. Bor.-Amer. 1: 188. (1832) Rydberg: Bull. Torrey Bot. Club 28: 174. (1901)
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