Potentilla bipinnatifida
Potentilla modesta
bipinnate cinquefoil, potentille bipinnatifide, tansy cinquefoil
modest cinquefoil
usually not sheathed with marcescent whole leaves.
ascending to erect, (1–)2–5 dm.
ascending to nearly erect, (0.3–)0.5–1.5(–2.5) dm.
subpinnate to subpalmate, (6–)10–25 cm;
petiole (2–)5–15 cm, long hairs dense, appressed, 1–2 mm, soft to ± stiff, short hairs absent, crisped hairs sparse, glands absent, sparse, or obscured;
leaflets 2–3 per side, on distal 1/6–1/3(–1/2) of leaf axis, separate to ± overlapping, terminal ones oblanceolate, (2–)3–6(–10) × 1–2(–3.5) cm, margins revolute, incised 3/4+ to midvein, undivided medial blade 1.5–6 mm wide, teeth 5–8 per side, ± linear, surfaces ± to strongly dissimilar, abaxial usually white, rarely grayish, long hairs abundant especially on veins, 1–2 mm, ± weak, short hairs absent or obscured, cottony (and crisped) hairs ± dense, glands absent or obscured, adaxial green to grayish, long hairs sparse to abundant, loosely appressed, 0.5–1.5 mm, short hairs absent or sparse, crisped and/or cottony hairs sparse to common, glands sparse to common.
usually palmate, rarely also ternate on same plant or subpalmate, 2–6(–8) cm;
petiole 0.5–3.5(–5) cm, long hairs common to abundant, ± ascending to almost spreading, 1–2 mm, weak to ± stiff, scarcely to ± verrucose, short and/or crisped hairs common to abundant, cottony hairs absent, glands sparse to common;
leaflets (3–)5, proximalmost separated by 0–1 mm, central obovate to oblanceolate-elliptic, 0.5–2(–2.5) × 0.5–1(–1.5) cm, petiolules 1 mm, distal 3/4 to nearly whole margin incised 1/2–3/4 to midvein, teeth 2–5 per side, (1–)2–5 mm, apical tufts 0.5–1 mm, abaxial surfaces white, long hairs common to abundant, cottony-crisped hairs dense, short hairs and glands absent or obscured, adaxial grayish green, long hairs common to abundant, 0.5–1.5 mm, ± stiff, rarely soft, short and/or crisped hairs sparse to common, cottony hairs absent, glands sparse to common.
2–4.
1–2.
(4–)10–50(–100)-flowered, congested or elongating in fruit.
(1–)3–10-flowered, congested or ± elongating in fruit, branch angle 20–35°.
0.2–0.8 cm (proximal to 2 cm).
0.2–0.5 cm, proximal to 1.2 cm.
epicalyx bractlets narrowly elliptic to lanceolate, 2.5–6 mm, lengths ± 2/3 times sepals, margins flat;
hypanthium 3–5 mm diam.;
sepals 3–6 mm, apex ± acute, abaxial surfaces: venation indistinct, glands absent, sparse, or obscured;
petals yellow, 3–5 × 3–4 mm, lengths ± equal to sepals;
filaments 0.5–2 mm, anthers 0.3–0.5 mm;
carpels 50–80, styles papillate-swollen in proximal 1/2–3/4+, 1–1.2 mm.
epicalyx bractlets narrowly elliptic, 1.5–3 × 0.5–1 mm;
hypanthium 3–4 mm diam.;
sepals 2.5–4 mm, apex ± acute, glands abundant, usually not obscured;
petals yellow, ± overlapping, 3.5–5 × 3–4 mm, slightly longer than sepals;
filaments 1 mm, anthers 0.3–0.7 mm;
carpels 20–40, styles 1 mm.
1–1.2 mm, smooth to faintly rugose.
1 mm.
= 56.
Potentilla bipinnatifida
Potentilla modesta
Potentilla bipinnatifida is similar to P. litoralis in habit and leaf dissection but has flat, silky epicalyx bractlets and sepals with no evident glands. Vestiture is generally silkier, and the silvery to bicolor leaves are white-cottony abaxially. The two species are sympatric in the plains of Canada, with some intergradation; P. bipinnatifida is also common south to Colorado, where it is found in intermontane meadows and sagebrush flats. Outlying populations occur in Blaine and Custer counties, Idaho, and Duchesne and Piute counties, Utah. Eastern collections from disturbed sites might be adventive.
Potentilla missourica Hornemann ex Lindley and P. normalis Besser ex Sprengel are older names for this species; both were rejected against a conserved P. bipinnatifida with designated lectotypes (see J. Soják 2008b).
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Potentilla modesta is the dominant component of what has previously been called P. rubricaulis in the Intermountain Region (for example, N. H. Holmgren 1997b, including illustration). This and other species of alpine Potentilla often grow in mixed populations, which contributes to confusion and mistaken identifications. In general, P. modesta has a more compact inflorescence and more consistently palmate basal leaves than sympatric members of the section. Intergrades with P. jepsonii (sect. Pensylvanicae) can also be problematic, though leaves of the latter are generally subpinnate and abaxially grayish rather than white.
The epithet modesta is misapplied in the combination Potentilla concinna var. modesta, which S. L. Welsh et al. (1993) used for plants mostly placed here in P. concinna var. divisa (sect. Concinnae).
(Discussion copyrighted by Flora of North America; reprinted with permission.)
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