Potentilla bipinnatifida |
Potentilla bimundorum |
|
---|---|---|
bipinnate cinquefoil, potentille bipinnatifide, tansy cinquefoil |
cutleaf cinquefoil, potentille des deux mondes, staghorn cinquefoil |
|
Stems | ascending to erect, (1–)2–5 dm. |
decumbent to ascending, rarely erect, 1–5 dm. |
Basal leaves | subpinnate to subpalmate, (6–)10–25 cm; petiole (2–)5–15 cm, long hairs dense, appressed, 1–2 mm, soft to ± stiff, short hairs absent, crisped hairs sparse, glands absent, sparse, or obscured; leaflets 2–3 per side, on distal 1/6–1/3(–1/2) of leaf axis, separate to ± overlapping, terminal ones oblanceolate, (2–)3–6(–10) × 1–2(–3.5) cm, margins revolute, incised 3/4+ to midvein, undivided medial blade 1.5–6 mm wide, teeth 5–8 per side, ± linear, surfaces ± to strongly dissimilar, abaxial usually white, rarely grayish, long hairs abundant especially on veins, 1–2 mm, ± weak, short hairs absent or obscured, cottony (and crisped) hairs ± dense, glands absent or obscured, adaxial green to grayish, long hairs sparse to abundant, loosely appressed, 0.5–1.5 mm, short hairs absent or sparse, crisped and/or cottony hairs sparse to common, glands sparse to common. |
subpinnate, 7–22(–25) cm; petiole 3–12(–15) cm, long hairs ± abundant, appressed to ascending, 0.5–3 mm, stiff, short hairs absent, cottony or crisped hairs absent, glands absent or sparse; leaflets 3(–4) per side, on distal 1/5–1/3(–1/2) of leaf axis, overlapping or not, terminal ones oblong to broadly oblong, 1.5–5 × 0.7–2 cm, margins strongly revolute, incised nearly to midvein, undivided medial blade 1–3 mm wide, teeth (3–)4–5 per side, linear, rarely narrowly oblong to narrowly lanceolate, surfaces strongly dissimilar, abaxial white or grayish white, long hairs abundant, ± 1 mm, stiff verrucose on veins, sparse and soft between veins, short hairs absent, crisped and cottony hairs dense, glands absent, adaxial dark green, long hairs sparse to common, tightly appressed, 0.5–1 mm, short hairs absent, cottony and crisped hairs absent, glands absent or sparse. |
Cauline leaves | 2–4. |
(1–)2–3. |
Inflorescences | (4–)10–50(–100)-flowered, congested or elongating in fruit. |
(5–)10–40+-flowered, ± congested to open. |
Pedicels | 0.2–0.8 cm (proximal to 2 cm). |
0.3–1 cm (proximal to 3 cm). |
Flowers | epicalyx bractlets narrowly elliptic to lanceolate, 2.5–6 mm, lengths ± 2/3 times sepals, margins flat; hypanthium 3–5 mm diam.; sepals 3–6 mm, apex ± acute, abaxial surfaces: venation indistinct, glands absent, sparse, or obscured; petals yellow, 3–5 × 3–4 mm, lengths ± equal to sepals; filaments 0.5–2 mm, anthers 0.3–0.5 mm; carpels 50–80, styles papillate-swollen in proximal 1/2–3/4+, 1–1.2 mm. |
epicalyx bractlets linear to narrowly oblong-lanceolate, 3–5 × 0.9–1.3 mm, lengths ± 3/4 times sepals, margins flat; hypanthium 2.5–3.5 mm diam.; sepals 4–7(–8) mm, apex acute to obtuse, abaxial surfaces: venation prominent at least proximally, glands ± sparse, obscured to evident; petals pale yellow, 3.5–5 × 3–5 mm, lengths ± equal to sepals; filaments 0.5–1.5 mm, anthers 0.5–0.6 mm; carpels 40–60, styles papillate-swollen at very base, if at all, 0.8–0.9 mm. |
Achenes | 1–1.2 mm, smooth to faintly rugose. |
1.3–1.5 mm, rugose. |
2n | = 56. |
= 28. |
Potentilla bipinnatifida |
Potentilla bimundorum |
|
Phenology | Flowering summer. | Flowering summer. |
Habitat | Open shortgrass prairie, alkaline bottoms, streamsides in sagebrush, disturbed sites | Road and railway verges, gravelly ruderal sites, acidic rocky outcrops, stream banks and gravel bars, sandy-gravelly flood plains, gravel ridges, sandy lakeshores, sandy steppe bluffs |
Elevation | 10–3400 m (0–11200 ft) | 0–1000 m (0–3300 ft) |
Distribution |
AK; CO; ID; MI; MN; MT; ND; NE; NM; SD; UT; WY; AB; MB; NT; ON; QC; SK; YT
|
AK; AB; MB; NT; NU; ON; QC; SK; YT; e Asia (Russian Far East) |
Discussion | Potentilla bipinnatifida is similar to P. litoralis in habit and leaf dissection but has flat, silky epicalyx bractlets and sepals with no evident glands. Vestiture is generally silkier, and the silvery to bicolor leaves are white-cottony abaxially. The two species are sympatric in the plains of Canada, with some intergradation; P. bipinnatifida is also common south to Colorado, where it is found in intermontane meadows and sagebrush flats. Outlying populations occur in Blaine and Custer counties, Idaho, and Duchesne and Piute counties, Utah. Eastern collections from disturbed sites might be adventive. Potentilla missourica Hornemann ex Lindley and P. normalis Besser ex Sprengel are older names for this species; both were rejected against a conserved P. bipinnatifida with designated lectotypes (see J. Soják 2008b). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Potentilla bimundorum is the correct name for what has been treated as P. multifida Linnaeus in North America. It is the only American representative of the mainly Eurasian P. multifida group, which is sometimes treated as a distinct section. J. Soják (2005) presented a useful survey and key to this group, showing that its species, in spite of being merged within P. multifida in the broad sense by many authors, are each distinct and with their own range. Potentilla multifida in the narrow sense is restricted to eastern Europe and western Siberia. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 9, p. 217. | FNA vol. 9, p. 218. |
Parent taxa | Rosaceae > subfam. Rosoideae > tribe Potentilleae > Potentilla > sect. Pensylvanicae | Rosaceae > subfam. Rosoideae > tribe Potentilleae > Potentilla > sect. Pensylvanicae |
Sibling taxa | ||
Synonyms | P. pensylvanica var. bipinnatifida | P. multifida subsp. bimundorum |
Name authority | Douglas: in W. J. Hooker, Fl. Bor.-Amer. 1: 188. (1832) | Soják: Č as. Nár. Muz. Praze Rada P ř ír. 141: 195. (1974) |
Web links |