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bipinnate cinquefoil, potentille bipinnatifide, tansy cinquefoil

Black Rock potentilla, Soldier Meadow or Black Rock cinquefoil, soldier meadows cinquefoil

Habit Plants ± rosetted; taproots fleshy-thickened.
Stems

ascending to erect, (1–)2–5 dm.

± prostrate, sometimes ascending in supporting vegetation, 1.5–5 dm, lengths 2–3 times basal leaves.

Basal leaves

subpinnate to subpalmate, (6–)10–25 cm;

petiole (2–)5–15 cm, long hairs dense, appressed, 1–2 mm, soft to ± stiff, short hairs absent, crisped hairs sparse, glands absent, sparse, or obscured;

leaflets 2–3 per side, on distal 1/6–1/3(–1/2) of leaf axis, separate to ± overlapping, terminal ones oblanceolate, (2–)3–6(–10) × 1–2(–3.5) cm, margins revolute, incised 3/4+ to midvein, undivided medial blade 1.5–6 mm wide, teeth 5–8 per side, ± linear, surfaces ± to strongly dissimilar, abaxial usually white, rarely grayish, long hairs abundant especially on veins, 1–2 mm, ± weak, short hairs absent or obscured, cottony (and crisped) hairs ± dense, glands absent or obscured, adaxial green to grayish, long hairs sparse to abundant, loosely appressed, 0.5–1.5 mm, short hairs absent or sparse, crisped and/or cottony hairs sparse to common, glands sparse to common.

pinnate with distal leaflets ± confluent, 5–12(–18) × 1–1.5(–2) cm;

petiole 0.5–1(–1.5) cm, straight hairs absent, cottony hairs absent, glands absent or sparse;

primary lateral leaflets (5–)10–15(–20) per side, on nearly whole leaf axis, ± overlapping, largest ones (or leaflet lobes) elliptic, 0.4–1(–2) × 0.2–0.5(–1) cm, simple and entire or 2(–3)-fid to base (sometimes shallowly toothed as well), segments 1–2(–3), narrowly to broadly elliptic, 4–10(–20) × (1–)2–3 mm, apical tufts absent or less than 0.5 mm, surfaces green, glaucous, straight hairs absent or sparse to common, tightly appressed, 0.5 mm or less, stiff, cottony hairs absent, glands absent or sparse.

Cauline leaves

2–4.

2–4.

Inflorescences

(4–)10–50(–100)-flowered, congested or elongating in fruit.

(5–)10–20-flowered, loosely cymose.

Pedicels

0.2–0.8 cm (proximal to 2 cm).

0.8–3(–4) cm, straight to ± recurved in fruit.

Flowers

epicalyx bractlets narrowly elliptic to lanceolate, 2.5–6 mm, lengths ± 2/3 times sepals, margins flat;

hypanthium 3–5 mm diam.;

sepals 3–6 mm, apex ± acute, abaxial surfaces: venation indistinct, glands absent, sparse, or obscured;

petals yellow, 3–5 × 3–4 mm, lengths ± equal to sepals;

filaments 0.5–2 mm, anthers 0.3–0.5 mm;

carpels 50–80, styles papillate-swollen in proximal 1/2–3/4+, 1–1.2 mm.

epicalyx bractlets lanceolate-elliptic to ovate-elliptic, 1.5–2.5(–3) × 1 mm;

hypanthium 2–5 mm diam.;

sepals (3–)4–5(–6) mm, apex acute to acuminate;

petals (3.5–)4.5–6.5 × 2.5–4 mm;

filaments 1.5–3 mm, anthers 0.6–0.8 mm;

carpels 3–10, styles 2–2.5 mm.

Achenes

1–1.2 mm, smooth to faintly rugose.

1.8–2.2 mm, smooth to ± rugose, not carunculate.

2n

= 56.

Potentilla bipinnatifida

Potentilla basaltica

Phenology Flowering summer. Flowering late spring–summer.
Habitat Open shortgrass prairie, alkaline bottoms, streamsides in sagebrush, disturbed sites Moist, subalkaline meadows in shrub steppe
Elevation 10–3400 m (0–11200 ft) 1300–1600 m (4300–5200 ft)
Distribution
from FNA
AK; CO; ID; MI; MN; MT; ND; NE; NM; SD; UT; WY; AB; MB; NT; ON; QC; SK; YT
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; NV
[BONAP county map]
Discussion

Potentilla bipinnatifida is similar to P. litoralis in habit and leaf dissection but has flat, silky epicalyx bractlets and sepals with no evident glands. Vestiture is generally silkier, and the silvery to bicolor leaves are white-cottony abaxially. The two species are sympatric in the plains of Canada, with some intergradation; P. bipinnatifida is also common south to Colorado, where it is found in intermontane meadows and sagebrush flats. Outlying populations occur in Blaine and Custer counties, Idaho, and Duchesne and Piute counties, Utah. Eastern collections from disturbed sites might be adventive.

Potentilla missourica Hornemann ex Lindley and P. normalis Besser ex Sprengel are older names for this species; both were rejected against a conserved P. bipinnatifida with designated lectotypes (see J. Soják 2008b).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Of conservation concern.

Potentilla basaltica is very distinctive in its glaucous, highly divided cylindric leaves with leaflets arranged in four ranks. The leaves strongly resemble those of some species of Ivesia, especially I. kingii S. Watson, but molecular analysis (B. Ertter et al. 1998) confirms a relationship with other members of sect. Multijugae.

Potentilla basaltica is known from only two localities, one in Lassen County, California, and the other in Humboldt County, Nevada. Both populations are highly localized, and P. basaltica is a candidate for federal listing as well as of conservation concern in both states.

The epithet basaltica was chosen as a reference to the Black Rock Desert, not habitat.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 9, p. 217. FNA vol. 9, p. 176.
Parent taxa Rosaceae > subfam. Rosoideae > tribe Potentilleae > Potentilla > sect. Pensylvanicae Rosaceae > subfam. Rosoideae > tribe Potentilleae > Potentilla > sect. Multijugae
Sibling taxa
P. albiflora, P. ambigens, P. anachoretica, P. angelliae, P. anglica, P. anserina, P. arenosa, P. argentea, P. arizonica, P. basaltica, P. bicrenata, P. biennis, P. biflora, P. bimundorum, P. brevifolia, P. breweri, P. bruceae, P. brunnescens, P. canadensis, P. concinna, P. cottamii, P. crantzii, P. crebridens, P. crinita, P. cristae, P. demotica, P. drummondii, P. effusa, P. elegans, P. erecta, P. flabellifolia, P. fragiformis, P. furcata, P. glaucophylla, P. gracilis, P. grayi, P. hickmanii, P. hippiana, P. holmgrenii, P. hookeriana, P. hyparctica, P. inclinata, P. intermedia, P. jepsonii, P. johnstonii, P. lasiodonta, P. litoralis, P. macounii, P. millefolia, P. modesta, P. morefieldii, P. multijuga, P. multisecta, P. nana, P. newberryi, P. nivea, P. norvegica, P. ovina, P. paucijuga, P. pedersenii, P. pensylvanica, P. plattensis, P. pseudosericea, P. pulchella, P. pulcherrima, P. recta, P. reptans, P. rhyolitica, P. rimicola, P. rivalis, P. robbinsiana, P. rubella, P. rubricaulis, P. sanguinea, P. saximontana, P. sierrae-blancae, P. simplex, P. sterilis, P. stipularis, P. subgorodkovii, P. subjuga, P. subvahliana, P. subviscosa, P. supina, P. thurberi, P. thuringiaca, P. tikhomirovii, P. townsendii, P. uliginosa, P. uschakovii, P. vahliana, P. verna, P. versicolor, P. villosa, P. villosula, P. vulcanicola, P. wheeleri
P. albiflora, P. ambigens, P. anachoretica, P. angelliae, P. anglica, P. anserina, P. arenosa, P. argentea, P. arizonica, P. bicrenata, P. biennis, P. biflora, P. bimundorum, P. bipinnatifida, P. brevifolia, P. breweri, P. bruceae, P. brunnescens, P. canadensis, P. concinna, P. cottamii, P. crantzii, P. crebridens, P. crinita, P. cristae, P. demotica, P. drummondii, P. effusa, P. elegans, P. erecta, P. flabellifolia, P. fragiformis, P. furcata, P. glaucophylla, P. gracilis, P. grayi, P. hickmanii, P. hippiana, P. holmgrenii, P. hookeriana, P. hyparctica, P. inclinata, P. intermedia, P. jepsonii, P. johnstonii, P. lasiodonta, P. litoralis, P. macounii, P. millefolia, P. modesta, P. morefieldii, P. multijuga, P. multisecta, P. nana, P. newberryi, P. nivea, P. norvegica, P. ovina, P. paucijuga, P. pedersenii, P. pensylvanica, P. plattensis, P. pseudosericea, P. pulchella, P. pulcherrima, P. recta, P. reptans, P. rhyolitica, P. rimicola, P. rivalis, P. robbinsiana, P. rubella, P. rubricaulis, P. sanguinea, P. saximontana, P. sierrae-blancae, P. simplex, P. sterilis, P. stipularis, P. subgorodkovii, P. subjuga, P. subvahliana, P. subviscosa, P. supina, P. thurberi, P. thuringiaca, P. tikhomirovii, P. townsendii, P. uliginosa, P. uschakovii, P. vahliana, P. verna, P. versicolor, P. villosa, P. villosula, P. vulcanicola, P. wheeleri
Synonyms P. pensylvanica var. bipinnatifida
Name authority Douglas: in W. J. Hooker, Fl. Bor.-Amer. 1: 188. (1832) Tiehm & Ertter: Brittonia 36: 228, fig. 1. (1984)
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