Potentilla bipinnatifida |
Potentilla anglica |
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bipinnate cinquefoil, potentille bipinnatifide, tansy cinquefoil |
English cinquefoil, potentille d'angleterre, trailing cinquefoil, trailing tormentil, wood cinquefoil |
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Stems | ascending to erect, (1–)2–5 dm. |
soon becoming prostrate, ± flagelliform, usually openly branched, eventually rooting at some nodes, (0.3–)1.5–7+ dm. |
Basal leaves | subpinnate to subpalmate, (6–)10–25 cm; petiole (2–)5–15 cm, long hairs dense, appressed, 1–2 mm, soft to ± stiff, short hairs absent, crisped hairs sparse, glands absent, sparse, or obscured; leaflets 2–3 per side, on distal 1/6–1/3(–1/2) of leaf axis, separate to ± overlapping, terminal ones oblanceolate, (2–)3–6(–10) × 1–2(–3.5) cm, margins revolute, incised 3/4+ to midvein, undivided medial blade 1.5–6 mm wide, teeth 5–8 per side, ± linear, surfaces ± to strongly dissimilar, abaxial usually white, rarely grayish, long hairs abundant especially on veins, 1–2 mm, ± weak, short hairs absent or obscured, cottony (and crisped) hairs ± dense, glands absent or obscured, adaxial green to grayish, long hairs sparse to abundant, loosely appressed, 0.5–1.5 mm, short hairs absent or sparse, crisped and/or cottony hairs sparse to common, glands sparse to common. |
± persistent, ternate or palmate, 2–10(–12) cm; petiole 1–7(–8) cm, long hairs sparse to abundant, appressed, 0.5–1 mm, stiff, glands absent; leaflets 3–5, central ± obovate to cuneate, 1–3(–3.5) × 0.8–2(–2.5) cm, distal ± 1/2 of margin incised 1/4–1/3(–1/2) to midvein, teeth 2–4 per side, surfaces similar, green, sparsely to moderately hairy. |
Cauline leaves | 2–4. |
2–3(–4) proximal to 1st flowering or branching node, usually well expanded at anthesis, usually ternate, 2–6(–10) cm; petiole 0.3–4(–8) cm; leaflets (3–)5, ± resembling or narrower than those of basal leaves, narrowly cuneate, 1–2(–3.5) × 0.8–2(–2.5) cm, apex rounded to obtuse. |
Inflorescences | (4–)10–50(–100)-flowered, congested or elongating in fruit. |
solitary flowers at stolon nodes. |
Pedicels | 0.2–0.8 cm (proximal to 2 cm). |
(1–)3–10(–17) cm. |
Flowers | epicalyx bractlets narrowly elliptic to lanceolate, 2.5–6 mm, lengths ± 2/3 times sepals, margins flat; hypanthium 3–5 mm diam.; sepals 3–6 mm, apex ± acute, abaxial surfaces: venation indistinct, glands absent, sparse, or obscured; petals yellow, 3–5 × 3–4 mm, lengths ± equal to sepals; filaments 0.5–2 mm, anthers 0.3–0.5 mm; carpels 50–80, styles papillate-swollen in proximal 1/2–3/4+, 1–1.2 mm. |
4(–5)-merous; epicalyx bractlets narrowly elliptic to oblong or ovate, 3–4(–7) × 1–1.5 mm, smaller than to slightly larger than sepals; hypanthium 2–4 mm diam.; sepals (3–)4–6 mm, apex broadly acute or acuminate; petals 6–9 × 5–9 mm, apex ± retuse; stamens 15–20, filaments 0.8–1.2 mm, anthers 0.8–1.2 mm; carpels 20–50, styles 0.9–1.5 mm. |
Achenes | 1–1.2 mm, smooth to faintly rugose. |
1–1.5(–1.8) mm, ± smooth. |
Rootstocks | ± erect, stout to slender, 1–4 cm. |
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2n | = 56. |
= 28, 56 (Europe). |
Potentilla bipinnatifida |
Potentilla anglica |
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Phenology | Flowering summer. | Flowering May–Aug(–Nov). |
Habitat | Open shortgrass prairie, alkaline bottoms, streamsides in sagebrush, disturbed sites | Moist flats and slopes, mainly on acidic soil |
Elevation | 10–3400 m (0–11200 ft) | 0–1200 m (0–3900 ft) |
Distribution |
AK; CO; ID; MI; MN; MT; ND; NE; NM; SD; UT; WY; AB; MB; NT; ON; QC; SK; YT
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CA; CO; ME; NY; OR; PA; UT; WA; BC; NL; NS; QC; SPM; Europe [Introduced in North America; introduced also in Atlantic Islands (Azores, Madeira), Pacific Islands (New Zealand), Australia]
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Discussion | Potentilla bipinnatifida is similar to P. litoralis in habit and leaf dissection but has flat, silky epicalyx bractlets and sepals with no evident glands. Vestiture is generally silkier, and the silvery to bicolor leaves are white-cottony abaxially. The two species are sympatric in the plains of Canada, with some intergradation; P. bipinnatifida is also common south to Colorado, where it is found in intermontane meadows and sagebrush flats. Outlying populations occur in Blaine and Custer counties, Idaho, and Duchesne and Piute counties, Utah. Eastern collections from disturbed sites might be adventive. Potentilla missourica Hornemann ex Lindley and P. normalis Besser ex Sprengel are older names for this species; both were rejected against a conserved P. bipinnatifida with designated lectotypes (see J. Soják 2008b). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Potentilla anglica apparently is a product of hybridization involving P. erecta and P. reptans (B. Matfield and J. R. Ellis 1972) that has become stabilized and distinct from both parents. In Europe, it forms back-cross hybrids with both P. erecta (P. ×suberecta Zimmeter) and P. reptans (P. ×mixta Nolte); these hybrids are not known from North America. A garden hybrid supposedly between P. anglica and P. nepalensis Hooker (known as P. ×tonguei Mallett) was found in Allegany State Park, New York, but this was likely cultivated rather than naturalized. Potentilla anglica probably was introduced even in Newfoundland (A. Kurtto et al. in J. Jalas et al. 1972+, vol. 13), contrary to the view expressed by M. L. Fernald (1950). The name Potentilla procumbens Sibthorp was previously used for this species; that is a superfluous and illegitimate name. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 9, p. 217. | FNA vol. 9, p. 135. |
Parent taxa | Rosaceae > subfam. Rosoideae > tribe Potentilleae > Potentilla > sect. Pensylvanicae | Rosaceae > subfam. Rosoideae > tribe Potentilleae > Potentilla > sect. Potentilla |
Sibling taxa | ||
Synonyms | P. pensylvanica var. bipinnatifida | |
Name authority | Douglas: in W. J. Hooker, Fl. Bor.-Amer. 1: 188. (1832) | Laicharding: Veg. Europ. 1: 475. (1790) |
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