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biennial cinquefoil, biennial or Greene's cinquefoil, Greene's cinquefoil

Morefield's cinquefoil

Stems

ascending to erect, (1–)2–6(–8.5) dm, hairs at base not stiff, not tubercle-based, glands sparse to abundant, conspicuous (to 1 mm, septate).

0.4–1.5(–1.7) dm, lengths 2–3 times basal leaves.

Leaves

ternate, (2–)4–12(–25) cm;

petiole (1–)2–8(–20) cm, long hairs sparse to common, spreading to ascending, 0.5–1.5(–3) mm, weak, crisped hairs usually absent, glands sparse to abundant, conspicuous (to 1 mm, septate);

leaflets 3, at tip of leaf axis, usually overlapping, largest ones mostly obovate or oval to nearly round, (0.5–)1–3(–4) × 0.3–3(–3.5) cm, distal 3/4 to ± whole margin evenly to unevenly incised 1/4–1/3(–1/2) to midvein, teeth 3–8(–13) per side, surfaces sparsely to moderately hairy, rarely glabrate, glands sparse to abundant at least abaxially.

Basal leaves

pinnate, usually at least 2 pairs of leaflets separate from terminal leaflets, distal leaflets usually distinct, 2–6 cm;

petiole 0.2–1.5 cm, straight hairs abundant, ± appressed, 1–1.5 mm, stiff, cottony hairs absent, glands absent or sparse;

leaflets 5–9, on distal (1/4–)1/2–3/4+ of leaf axis, overlapping, proximal pair separated from others by 2–10 mm of leaf axis, central leaflets oblanceolate to narrowly obovate, (0.5–)1–2(–2.5) × 0.5–1.2 cm, petiolules 0–1 mm, distal 3/4 of margins incised 3/4 to midvein (sometimes medially cleft as well), teeth 3–4 per side, separate to slightly overlapping, (1–)2–5 mm, surfaces ± to strongly dissimilar, abaxial grayish to white, straight hairs ± abundant, ± appressed, 1 mm, weak to stiff (especially on veins), cottony hairs abundant to dense, glands absent or obscured, adaxial greenish to grayish, straight hairs ± abundant, appressed, 0.5–1.5 mm, stiff, cottony hairs absent, rarely sparse, glands sparse.

Inflorescences

(10–)20–70+-flowered.

3–15-flowered.

Pedicels

0.2–1(–2) cm.

1–2.5(–3) cm.

Flowers

epicalyx bractlets narrowly oblanceolate to narrowly obovate or elliptic, (1.5–)2–3.5(–4.5) × 0.5–1.5(–2) mm;

hypanthium (2–)3–4(–5.5) mm diam.;

sepals (2–)3–5 mm, apex broadly acute to obtuse;

petals pale yellow to yellow, broadly oblanceolate to obovate, (1–)2–2.7 × 1.5 mm;

stamens 10 or 15, filaments 0.5–1.2 mm, anthers 0.2–0.3 mm;

carpels 40–100, styles 0.5–0.7 mm.

epicalyx bractlets ± ovate-elliptic, 2–4 × 0.8–2 mm;

hypanthium 2.5–5 mm diam.;

sepals 3.5–5 mm, apex ± acute;

petals 4–6 × 3–5 mm;

filaments 1–2 mm, anthers 0.5–1 mm;

carpels 15–20, styles 2 mm.

Achenes

whitish or pale yellowish, 0.5–0.9 mm, smooth, without a corky protuberance.

1.5–2 mm, smooth to faintly rugose.

Potentilla biennis

Potentilla morefieldii

Phenology Flowering summer. Flowering summer.
Habitat Moist meadows, stream banks, ditches, seepages Alpine tundra and fellfields, mostly on dolomite substrates
Elevation 400–3100 m (1300–10200 ft) 3500–4000 m (11500–13100 ft)
Distribution
from FNA
AK; AZ; CA; CO; ID; MT; ND; NM; NV; OR; UT; WA; WY; AB; BC; SK; YT
[WildflowerSearch map]
[BONAP county map]
from FNA
CA
[BONAP county map]
Discussion

Potentilla biennis may be recognized by its septate hairs with a round, terminal gland, which dominate the vestiture especially at the base of the plant. Stems are more commonly erect and leaflets rounder than those of P. rivalis. The species is most common along the eastern side of the Sierra Nevada of California, across the Intermountain Region to the Rocky Mountains. All specimens seen by the authors from Baja California, Mexico, which were previously referred to P. biennis are now assigned to P. rivalis.

J. Soják (1996) lectotypified Potentilla millegrana Engelmann ex Lehmann upon a collection of P. rivalis, thus preventing P. biennis from being a later taxonomic synonym.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Of conservation concern.

Potentilla morefieldii is known only from the White Mountains and adjacent Sierra Nevada. Plants with similar aspect but lacking strigose vestiture occur in White Pine County, Nevada; their disposition is uncertain. Potentilla morefieldii is sometimes confused with P. breweri (sect. Multijugae), but the latter differs in having more or less similar leaflet surfaces with cottony hairs nearly as abundant adaxially as abaxially. Potentilla breweri also lacks the stiff adaxial leaflet hairs that characterize most species in sect. Concinnae, including P. morefieldii.

W. L. Jepson (1909–1943, vol. 2) misapplied the name Potentilla pseudosericea var. grandiflora Th. Wolf, which is a synonym of an unresolved species from the Rocky Mountains (B. Ertter 1992; Ertter et al. 2013), to P. morefieldii.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 9, p. 140. FNA vol. 9, p. 181.
Parent taxa Rosaceae > subfam. Rosoideae > tribe Potentilleae > Potentilla > sect. Rivales Rosaceae > subfam. Rosoideae > tribe Potentilleae > Potentilla > sect. Concinnae
Sibling taxa
P. albiflora, P. ambigens, P. anachoretica, P. angelliae, P. anglica, P. anserina, P. arenosa, P. argentea, P. arizonica, P. basaltica, P. bicrenata, P. biflora, P. bimundorum, P. bipinnatifida, P. brevifolia, P. breweri, P. bruceae, P. brunnescens, P. canadensis, P. concinna, P. cottamii, P. crantzii, P. crebridens, P. crinita, P. cristae, P. demotica, P. drummondii, P. effusa, P. elegans, P. erecta, P. flabellifolia, P. fragiformis, P. furcata, P. glaucophylla, P. gracilis, P. grayi, P. hickmanii, P. hippiana, P. holmgrenii, P. hookeriana, P. hyparctica, P. inclinata, P. intermedia, P. jepsonii, P. johnstonii, P. lasiodonta, P. litoralis, P. macounii, P. millefolia, P. modesta, P. morefieldii, P. multijuga, P. multisecta, P. nana, P. newberryi, P. nivea, P. norvegica, P. ovina, P. paucijuga, P. pedersenii, P. pensylvanica, P. plattensis, P. pseudosericea, P. pulchella, P. pulcherrima, P. recta, P. reptans, P. rhyolitica, P. rimicola, P. rivalis, P. robbinsiana, P. rubella, P. rubricaulis, P. sanguinea, P. saximontana, P. sierrae-blancae, P. simplex, P. sterilis, P. stipularis, P. subgorodkovii, P. subjuga, P. subvahliana, P. subviscosa, P. supina, P. thurberi, P. thuringiaca, P. tikhomirovii, P. townsendii, P. uliginosa, P. uschakovii, P. vahliana, P. verna, P. versicolor, P. villosa, P. villosula, P. vulcanicola, P. wheeleri
P. albiflora, P. ambigens, P. anachoretica, P. angelliae, P. anglica, P. anserina, P. arenosa, P. argentea, P. arizonica, P. basaltica, P. bicrenata, P. biennis, P. biflora, P. bimundorum, P. bipinnatifida, P. brevifolia, P. breweri, P. bruceae, P. brunnescens, P. canadensis, P. concinna, P. cottamii, P. crantzii, P. crebridens, P. crinita, P. cristae, P. demotica, P. drummondii, P. effusa, P. elegans, P. erecta, P. flabellifolia, P. fragiformis, P. furcata, P. glaucophylla, P. gracilis, P. grayi, P. hickmanii, P. hippiana, P. holmgrenii, P. hookeriana, P. hyparctica, P. inclinata, P. intermedia, P. jepsonii, P. johnstonii, P. lasiodonta, P. litoralis, P. macounii, P. millefolia, P. modesta, P. multijuga, P. multisecta, P. nana, P. newberryi, P. nivea, P. norvegica, P. ovina, P. paucijuga, P. pedersenii, P. pensylvanica, P. plattensis, P. pseudosericea, P. pulchella, P. pulcherrima, P. recta, P. reptans, P. rhyolitica, P. rimicola, P. rivalis, P. robbinsiana, P. rubella, P. rubricaulis, P. sanguinea, P. saximontana, P. sierrae-blancae, P. simplex, P. sterilis, P. stipularis, P. subgorodkovii, P. subjuga, P. subvahliana, P. subviscosa, P. supina, P. thurberi, P. thuringiaca, P. tikhomirovii, P. townsendii, P. uliginosa, P. uschakovii, P. vahliana, P. verna, P. versicolor, P. villosa, P. villosula, P. vulcanicola, P. wheeleri
Name authority Greene: Fl. Francisc., 65. (1891) Ertter: Brittonia 44: 432, fig. 1. (1992)
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