Potentilla biennis |
Potentilla cristae |
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biennial cinquefoil, biennial or Greene's cinquefoil, Greene's cinquefoil |
crested cinquefoil, crested potentilla |
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Habit | Plants tufted to densely matted; caudex branches usually short, stout, sometimes embedded in old leaf bases. | |
Stems | ascending to erect, (1–)2–6(–8.5) dm, hairs at base not stiff, not tubercle-based, glands sparse to abundant, conspicuous (to 1 mm, septate). |
ascending to erect, 0.3–2 dm, lengths 2–3 times basal leaves. |
Leaves | ternate, (2–)4–12(–25) cm; petiole (1–)2–8(–20) cm, long hairs sparse to common, spreading to ascending, 0.5–1.5(–3) mm, weak, crisped hairs usually absent, glands sparse to abundant, conspicuous (to 1 mm, septate); leaflets 3, at tip of leaf axis, usually overlapping, largest ones mostly obovate or oval to nearly round, (0.5–)1–3(–4) × 0.3–3(–3.5) cm, distal 3/4 to ± whole margin evenly to unevenly incised 1/4–1/3(–1/2) to midvein, teeth 3–8(–13) per side, surfaces sparsely to moderately hairy, rarely glabrate, glands sparse to abundant at least abaxially. |
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Basal leaves | not in ranks, ternate, 1–5(–9) cm; stipules: apex acute to obtuse; petiole 1–4 cm, long hairs absent or sparse to common, spreading to ascending, 0.5–2 mm, ± weak, glands sparse to abundant; leaflets 3, central flabellate, 0.4–2 × 0.5–1.5 cm, petiolule 1–3 mm, margins flat, distal 3/4+ deeply 3–5-lobed (sinuses extending 1/2 to nearly to midvein), lobes unevenly incised 1/4–1/2 to midvein, teeth 2–5 per lobe, surfaces similar, dark green, hairs ± sparse, 0.5–1.5 mm, glands sparse to abundant. |
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Inflorescences | (10–)20–70+-flowered. |
1–7-flowered. |
Pedicels | 0.2–1(–2) cm. |
straight, 0.2–1.2(–2) cm, not much longer in fruit than in flower. |
Flowers | epicalyx bractlets narrowly oblanceolate to narrowly obovate or elliptic, (1.5–)2–3.5(–4.5) × 0.5–1.5(–2) mm; hypanthium (2–)3–4(–5.5) mm diam.; sepals (2–)3–5 mm, apex broadly acute to obtuse; petals pale yellow to yellow, broadly oblanceolate to obovate, (1–)2–2.7 × 1.5 mm; stamens 10 or 15, filaments 0.5–1.2 mm, anthers 0.2–0.3 mm; carpels 40–100, styles 0.5–0.7 mm. |
epicalyx bractlets broadly ovate, 2–4 × 2 mm, margins flat; hypanthium 3.5–6 mm diam.; sepals 3–5 mm, apex broadly acute to obtuse; petals yellow, 3–5.5 × 4–5 mm; filaments 0.7–1.7 mm, anthers 0.5–0.6 mm; carpels 25–40, styles tapered-filiform, ± papillate-swollen in proximal 1/5–/1/3, 1.3–2 mm. |
Achenes | whitish or pale yellowish, 0.5–0.9 mm, smooth, without a corky protuberance. |
1–1.5 mm, dorsally crested. |
2n | = 42. |
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Potentilla biennis |
Potentilla cristae |
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Phenology | Flowering summer. | Flowering summer. |
Habitat | Moist meadows, stream banks, ditches, seepages | Rocky, open, serpentine slopes, in conifer woodlands |
Elevation | 400–3100 m (1300–10200 ft) | 1800–2800 m (5900–9200 ft) |
Distribution |
AK; AZ; CA; CO; ID; MT; ND; NM; NV; OR; UT; WA; WY; AB; BC; SK; YT
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CA |
Discussion | Potentilla biennis may be recognized by its septate hairs with a round, terminal gland, which dominate the vestiture especially at the base of the plant. Stems are more commonly erect and leaflets rounder than those of P. rivalis. The species is most common along the eastern side of the Sierra Nevada of California, across the Intermountain Region to the Rocky Mountains. All specimens seen by the authors from Baja California, Mexico, which were previously referred to P. biennis are now assigned to P. rivalis. J. Soják (1996) lectotypified Potentilla millegrana Engelmann ex Lehmann upon a collection of P. rivalis, thus preventing P. biennis from being a later taxonomic synonym. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. Potentilla cristae is known only from Cory Peak, Mount Eddy, and the Marble Mountains in northwestern California. In addition to the characteristics in the key, the species is distinctive in having a low dorsal crest on the achenes, hence the specific epithet. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 9, p. 140. | FNA vol. 9, p. 191. |
Parent taxa | Rosaceae > subfam. Rosoideae > tribe Potentilleae > Potentilla > sect. Rivales | Rosaceae > subfam. Rosoideae > tribe Potentilleae > Potentilla > sect. Aureae |
Sibling taxa | ||
Name authority | Greene: Fl. Francisc., 65. (1891) | Ferlatte & Strother: Madroño 37: 190, fig. 1. (1990) |
Web links |
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