Potentilla argentea |
Potentilla arenosa |
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hoary cinquefoil, potentille argentée, silver cinquefoil, silver-leaf cinquefoil, silvery cinquefoil |
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Habit | Plants scarcely to ± tufted. | |||||
Caudex branches | thick, not columnar, not sheathed with marcescent whole leaves. |
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Stems | mostly decumbent to ascending, 1–6 dm. |
ascending to erect, (0.3–)0.8–2.5(–4.5) dm, lengths (2–)3–5 times basal leaves. |
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Basal leaves | palmate. |
1.5–12(–20) cm; petiole 1–7(–15) cm, long hairs sparse to abundant, spreading to ± ascending, rarely loosely appressed, 1–2(–2.5) mm, usually stiff, sometimes weak (subsp. chamissonis), verrucose, short and/or crisped hairs absent or sparse to abundant, cottony hairs absent, glands absent or sparse; leaflets separate to ± overlapping, central obovate, 1–3.5(–4.5) × 0.5–2(–3) cm, usually petiolulate, petiolule to 5 mm, base cuneate, margins slightly revolute, distal ± 3/4 incised ± 1/2 to midvein, teeth (2–)3–4(–6) per side, ± approximate to distant, surfaces dissimilar, often strongly so, abaxial white to gray, long hairs 0.5–1.8 mm, cottony-crisped hairs ± dense, adaxial green, sometimes grayish green, long hairs sparse to abundant, short-crisped hairs sparse to abundant. |
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Cauline leaves | 2–9, proximal ones 2–7(–10) cm; proximal petioles 1–4(–7) cm, long hairs absent or sparse, ± ascending, 1–2 mm, soft, short and crisped hairs absent or obscured, cottony hairs dense, glands absent or obscured; leaflets 5(–7), central one ± oblanceolate, (0.5–)1–3 × 0.3–1(–1.3) cm, margins revolute, distal 1/2–2/3 evenly or unevenly incised 1/2–3/4+ to midvein, teeth 2–3 per side (more if lobed or secondarily toothed), surfaces strongly dissimilar, abaxial white, long hairs ± sparse (mostly on veins), 1–2 mm, weak, short and crisped hairs absent or obscured, cottony hairs dense, glands absent or obscured, adaxial long hairs absent or sparse to common, short or crisped hairs absent or sparse, cottony hairs absent, glands absent or sparse. |
(0–)1–2. |
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Inflorescence(s) | 10–80-flowered. |
1–7(–15)-flowered. |
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Pedicels | 0.3–1.5(–2) cm. |
1.5–5 cm in flower, to 6(–10) cm in fruit. |
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Flowers | epicalyx bractlets oblong to narrowly ovate, 1.5–3 × 0.7–1.2 mm, lengths 2/3–1 times sepals; sepals 2–4.5 mm, apex ± acute; petals (2–)2.5–4 × 1.5–3 mm; filaments 0.8–1.5 mm, anthers 0.3–0.6(–0.8) mm; carpels 30–60, styles 0.6–0.9 mm, often strongly papillate-swollen proximally. |
epicalyx bractlets linear-lanceolate to lanceolate-elliptic, 2–5(–7) × 0.4–1.2(–1.5) mm, 1/4–1/2 as wide as sepals, margins usually flat, red glands absent or sparse and inconspicuous; hypanthium 3–5 mm diam.; sepals 3–6(–8) mm, apex acute; petals 4–7(–10) × 4–7(–9) mm, ± longer than sepals; filaments 0.8–1 mm, anthers 0.4 mm; carpels 28–40, apical hairs absent, styles conic-columnar, strongly papillate-swollen in proximal 1/5–1/3, 1–1.5 mm. |
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Achenes | 0.8–1.1 mm, smooth to lightly rugose. |
1.1 mm. |
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2n | = 14, 28, 35, 42, 56, 62 (Eurasia). |
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Potentilla argentea |
Potentilla arenosa |
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Phenology | Flowering spring–summer. | |||||
Habitat | Dry flats and slopes, roadsides, dry ditches, other open sites, in grasslands, oak and conifer woodlands | |||||
Elevation | 0–2000 m (0–6600 ft) | |||||
Distribution |
CO; CT; DC; IA; ID; IL; IN; KY; MA; MD; ME; MI; MN; MO; MT; NC; ND; NH; NJ; NY; OH; OR; PA; RI; SD; TN; VA; VT; WA; WI; WV; WY; AB; BC; MB; NB; NF; NS; ON; PE; QC; SK; SPM; Eurasia [Introduced in North America; introduced also in Pacific Islands (New Zealand)]
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AK; AB; BC; MB; NT; NU; QC; SK; YT; Eurasia |
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Discussion | The bright, silvery vestiture of the leaves quickly distinguishes Potentilla argentea from P. inclinata and P. intermedia. Although treated here as introduced, some (P. A. Rydberg 1898; P. A. Werner and J. D. Soule 1976) considered P. argentea to be probably native in eastern North America. The Potentilla argentea species group is an amphi- or apomictic polyploid complex that has been the subject of numerous cytological, sexual, and molecular studies (for example, A. Müntzing and G. Müntzing 1945; A. Müntzing 1958; J. Paule et al. 2011). Taxonomic treatments have ranged from a highly polymorphic single species to an abundance of species, varieties, and forms (T. Wolf 1908). This treatment is at the conservative end of the spectrum. Some New England references (R. C. Bean et al. 1967; F. C. Seymour 1969) distinguish plants with densely white-hairy adaxial leaflet surfaces as var. pseudocalabra Th. Wolf, a name based on plants from southeastern Europe. Molecular analysis by Paule et al. supports the conclusion by Wolf that the variety consists of intermediates between P. argentea and P. calabra Tenore. The optimum taxonomic disposition of var. pseudocalabra, and whether any North American plants are properly included, remains to be determined. Potentilla argentea has been reported as an active agent in treating cirrhosis of the liver (N. T. Starostenko and V. N. Starostenko 1971). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Subspecies 2 (2 in the flora). The name Potentilla arenosa is now used for most arctic and subarctic plants previously treated as P. hookeriana or P. nivea subsp. hookeriana (Lehmann) Hiitonen. As noted by J. Soják (1986), the type of P. hookeriana has quinate leaves; that name is now restricted to a Rocky Mountain species in sect. Rubricaules. The arctic and subarctic material was briefly (1989–1999) called P. nivea, as discussed under that species. Because the type of P. nivea var. arenosa and other northern Asian specimens correspond closely to the North American plants, the name P. arenosa is assigned here. The two subspecies differ only in one character, the petiole hairs, but are largely allopatric. Subspecies arenosa occurs in western and northern Greenland, northern North America (very northern in the east), and northern Asia (and perhaps northeasternmost European Russia); subsp. chamissonis occurs in southern Greenland, northeastern North America (more southern than subsp. arenosa), and northern Europe at least east to the Urals. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 9, p. 144. | FNA vol. 9, p. 200. | ||||
Parent taxa | Rosaceae > subfam. Rosoideae > tribe Potentilleae > Potentilla > sect. Terminales | Rosaceae > subfam. Rosoideae > tribe Potentilleae > Potentilla > sect. Niveae | ||||
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Subordinate taxa | ||||||
Synonyms | P. nivea var. arenosa | |||||
Name authority | Linnaeus: Sp. Pl. 1: 497. (1753) | (Turczaninow) Juzepczuk: in V. L. Komarov et al., Fl. URSS 10: 137. (1941) | ||||
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