Potentilla argentea |
Potentilla ambigens |
|
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hoary cinquefoil, potentille argentée, silver cinquefoil, silver-leaf cinquefoil, silvery cinquefoil |
silkyleaf cinquefoil, silkyleaf or giant or southern Rocky Mountain cinquefoil |
|
Stems | mostly decumbent to ascending, 1–6 dm. |
(2–)3–7(–8) dm, lengths (1–)1.5–2.5 times basal leaves. |
Basal leaves | palmate. |
pinnate, 15–45(–50) cm; petiole (5–)8–20 cm, long hairs abundant to dense, ascending to appressed, (1–)2–3 mm, ± stiff, rarely weak, short or crisped hairs absent or sparse, cottony hairs absent, glands sparse, often obscured; leaflets not conduplicate, lateral ones evenly paired, (3–)4–7 per side on distal ± 1/2 of leaf axis, most pairs ± decurrent, rachis winged, distal sometimes confluent with terminal leaflet, larger leaflets narrowly obovate, (2–)3–7 × 1–2.5 cm, distal 3/4+ of margin incised 1/3 to midvein, teeth 6–18 per side, 1–4 mm, surfaces ± similar, abaxial grayish, long hairs sparse to common (mostly on veins), 0.5–2 mm, ± stiff, rarely weak, short-crisped hairs absent or sparse to common, cottony hairs absent, glands sparse to common, adaxial green to grayish, long and short hairs not differentiated, sparse to common, crisped and cottony hairs absent, glands absent or sparse. |
Cauline leaves | 2–9, proximal ones 2–7(–10) cm; proximal petioles 1–4(–7) cm, long hairs absent or sparse, ± ascending, 1–2 mm, soft, short and crisped hairs absent or obscured, cottony hairs dense, glands absent or obscured; leaflets 5(–7), central one ± oblanceolate, (0.5–)1–3 × 0.3–1(–1.3) cm, margins revolute, distal 1/2–2/3 evenly or unevenly incised 1/2–3/4+ to midvein, teeth 2–3 per side (more if lobed or secondarily toothed), surfaces strongly dissimilar, abaxial white, long hairs ± sparse (mostly on veins), 1–2 mm, weak, short and crisped hairs absent or obscured, cottony hairs dense, glands absent or obscured, adaxial long hairs absent or sparse to common, short or crisped hairs absent or sparse, cottony hairs absent, glands absent or sparse. |
(0–)1–3. |
Inflorescences | 10–80-flowered. |
(10–)20–60-flowered. |
Pedicels | 0.3–1.5(–2) cm. |
(0.5–)1–3(–6) cm. |
Flowers | epicalyx bractlets oblong to narrowly ovate, 1.5–3 × 0.7–1.2 mm, lengths 2/3–1 times sepals; sepals 2–4.5 mm, apex ± acute; petals (2–)2.5–4 × 1.5–3 mm; filaments 0.8–1.5 mm, anthers 0.3–0.6(–0.8) mm; carpels 30–60, styles 0.6–0.9 mm, often strongly papillate-swollen proximally. |
epicalyx bractlets lanceolate to narrowly ovate or elliptic, 3–6 × 1–1.5 mm, 2/3 as long as to nearly equal to sepals, abaxial vestiture ± similar to sepals, not glabrescent, straight hairs sparse to common, crisped or cottony hairs usually absent; hypanthium 3–6 mm diam.; sepals 4–7 mm, apex acute to acuminate; petals (6–)7–10 × 5–9 mm; filaments (1–)1.5–4 mm, anthers 0.7–0.9(–1) mm; carpels 20–30, styles (1–)1.8–3 mm. |
Achenes | 0.8–1.1 mm, smooth to lightly rugose. |
1.4–1.6 mm, smooth. |
2n | = 14, 28, 35, 42, 56, 62 (Eurasia). |
= 82. |
Potentilla argentea |
Potentilla ambigens |
|
Phenology | Flowering spring–summer. | Flowering summer. |
Habitat | Dry flats and slopes, roadsides, dry ditches, other open sites, in grasslands, oak and conifer woodlands | Dry meadows and adjacent slopes, in conifer woodlands |
Elevation | 0–2000 m (0–6600 ft) | 2000–2700 m (6600–8900 ft) |
Distribution |
CO; CT; DC; IA; ID; IL; IN; KY; MA; MD; ME; MI; MN; MO; MT; NC; ND; NH; NJ; NY; OH; OR; PA; RI; SD; TN; VA; VT; WA; WI; WV; WY; AB; BC; MB; NB; NF; NS; ON; PE; QC; SK; SPM; Eurasia [Introduced in North America; introduced also in Pacific Islands (New Zealand)]
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CO; NM; WY |
Discussion | The bright, silvery vestiture of the leaves quickly distinguishes Potentilla argentea from P. inclinata and P. intermedia. Although treated here as introduced, some (P. A. Rydberg 1898; P. A. Werner and J. D. Soule 1976) considered P. argentea to be probably native in eastern North America. The Potentilla argentea species group is an amphi- or apomictic polyploid complex that has been the subject of numerous cytological, sexual, and molecular studies (for example, A. Müntzing and G. Müntzing 1945; A. Müntzing 1958; J. Paule et al. 2011). Taxonomic treatments have ranged from a highly polymorphic single species to an abundance of species, varieties, and forms (T. Wolf 1908). This treatment is at the conservative end of the spectrum. Some New England references (R. C. Bean et al. 1967; F. C. Seymour 1969) distinguish plants with densely white-hairy adaxial leaflet surfaces as var. pseudocalabra Th. Wolf, a name based on plants from southeastern Europe. Molecular analysis by Paule et al. supports the conclusion by Wolf that the variety consists of intermediates between P. argentea and P. calabra Tenore. The optimum taxonomic disposition of var. pseudocalabra, and whether any North American plants are properly included, remains to be determined. Potentilla argentea has been reported as an active agent in treating cirrhosis of the liver (N. T. Starostenko and V. N. Starostenko 1971). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. Potentilla ambigens is one of the more distinctive species in sect. Leucophyllae, comprising exceptionally large plants with large, coarsely hairy leaves. Although sometimes confused with P. hippiana, P. ambigens lacks the crisped-cottony vestiture of that species and is usually morphologically distinctive even where both species occur together (D. G. Anderson, www.fs.fed.us/r2/projects/scp/assessments/potentillaambigens.pdf). Potentilla ambigens occurs in Wyoming (at least historically), along the eastern slope of the Rocky Mountains to northern Colorado, and in scattered locations in the high mountains of New Mexico as far south as the Sacramento Mountains, Otero County. The species is of conservation concern in Colorado and has not been documented in Wyoming since 1900 (D. G. Anderson, www.fs.fed.us/r2/projects/scp/assessments/potentillaambigens.pdf). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 9, p. 144. | FNA vol. 9, p. 164. |
Parent taxa | Rosaceae > subfam. Rosoideae > tribe Potentilleae > Potentilla > sect. Terminales | Rosaceae > subfam. Rosoideae > tribe Potentilleae > Potentilla > sect. Leucophyllae |
Sibling taxa | ||
Name authority | Linnaeus: Sp. Pl. 1: 497. (1753) | Greene: Erythea 1: 5. (1893) |
Web links |
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