Potentilla anglica |
Potentilla subvahliana |
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English cinquefoil, potentille d'angleterre, trailing cinquefoil, trailing tormentil, wood cinquefoil |
high arctic cinquefoil |
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Habit | Plants usually cushion-forming. | |
Caudex branches | stout, columnar, sheathed with marcescent whole leaves. |
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Stems | soon becoming prostrate, ± flagelliform, usually openly branched, eventually rooting at some nodes, (0.3–)1.5–7+ dm. |
erect, 0.2–0.6(–1.1) dm, lengths 2–4 times basal leaves. |
Basal leaves | ± persistent, ternate or palmate, 2–10(–12) cm; petiole 1–7(–8) cm, long hairs sparse to abundant, appressed, 0.5–1 mm, stiff, glands absent; leaflets 3–5, central ± obovate to cuneate, 1–3(–3.5) × 0.8–2(–2.5) cm, distal ± 1/2 of margin incised 1/4–1/3(–1/2) to midvein, teeth 2–4 per side, surfaces similar, green, sparsely to moderately hairy. |
0.5–2.5(–3) cm; petiole 0.3–1.5(–2) cm, long hairs common to abundant, spreading to ascending, 1(–2) mm, ± weak, rarely stiff, smooth, crisped/short-cottony hairs usually absent, sometimes sparse, glands sparse to common; leaflets separate to slightly overlapping, central obovate, 0.5–1.2(–1.5) × 0.4–0.9(–1) cm, subsessile to short-petiolulate, base cuneate, margins revolute, distal 1/2 incised 1/2–3/4 to midvein, teeth (1–)2–3(–4) per side, ± distant, surfaces ± dissimilar, abaxial grayish or yellowish white, long hairs 0.5–1 mm, cottony-crisped hairs ± dense (often obscured by long hairs), adaxial dark green to greenish gray, long hairs ± abundant, other hairs absent. |
Cauline leaves | 2–3(–4) proximal to 1st flowering or branching node, usually well expanded at anthesis, usually ternate, 2–6(–10) cm; petiole 0.3–4(–8) cm; leaflets (3–)5, ± resembling or narrower than those of basal leaves, narrowly cuneate, 1–2(–3.5) × 0.8–2(–2.5) cm, apex rounded to obtuse. |
(0–)1(–2). |
Inflorescences | solitary flowers at stolon nodes. |
1(–2)-flowered. |
Pedicels | (1–)3–10(–17) cm. |
1–2(–3) cm in flower, to 5.5 cm in fruit. |
Flowers | 4(–5)-merous; epicalyx bractlets narrowly elliptic to oblong or ovate, 3–4(–7) × 1–1.5 mm, smaller than to slightly larger than sepals; hypanthium 2–4 mm diam.; sepals (3–)4–6 mm, apex broadly acute or acuminate; petals 6–9 × 5–9 mm, apex ± retuse; stamens 15–20, filaments 0.8–1.2 mm, anthers 0.8–1.2 mm; carpels 20–50, styles 0.9–1.5 mm. |
epicalyx bractlets lanceolate to elliptic-ovate, rarely linear, 3–6(–7) × 0.8–2(–2.3) mm, (1/2–)2/3 to nearly as wide as sepals, margins flat or ± revolute, red glands absent; hypanthium 2.5–4 mm diam.; sepals 3–5(–6) mm, apex obtuse to subacute; petals 4–8(–9) × 4–9 mm, longer than sepals; filaments 1–1.3 mm, anthers 0.3–0.5 mm; carpels 25–35, apical hairs absent, styles conic-columnar, ± papillate-swollen on less than proximal 1/5, 0.9–1.1 mm. |
Achenes | 1–1.5(–1.8) mm, ± smooth. |
1.2–1.6 mm. |
Rootstocks | ± erect, stout to slender, 1–4 cm. |
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2n | = 28, 56 (Europe). |
= 28 (Russian Far East). |
Potentilla anglica |
Potentilla subvahliana |
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Phenology | Flowering May–Aug(–Nov). | Flowering late spring to summer. |
Habitat | Moist flats and slopes, mainly on acidic soil | Rocky alpine heaths, rock outcrops and crevices, scree and talus, dry tundra, coastal bluffs, stabilized sand dunes, usually on calcareous bedrock |
Elevation | 0–1200 m (0–3900 ft) | 0–1700 m (0–5600 ft) |
Distribution |
CA; CO; ME; NY; OR; PA; UT; WA; BC; NL; NS; QC; SPM; Europe [Introduced in North America; introduced also in Atlantic Islands (Azores, Madeira), Pacific Islands (New Zealand), Australia]
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AK; AB; BC; NT; NU; QC; YT; Greenland; e Asia (Russian Far East) |
Discussion | Potentilla anglica apparently is a product of hybridization involving P. erecta and P. reptans (B. Matfield and J. R. Ellis 1972) that has become stabilized and distinct from both parents. In Europe, it forms back-cross hybrids with both P. erecta (P. ×suberecta Zimmeter) and P. reptans (P. ×mixta Nolte); these hybrids are not known from North America. A garden hybrid supposedly between P. anglica and P. nepalensis Hooker (known as P. ×tonguei Mallett) was found in Allegany State Park, New York, but this was likely cultivated rather than naturalized. Potentilla anglica probably was introduced even in Newfoundland (A. Kurtto et al. in J. Jalas et al. 1972+, vol. 13), contrary to the view expressed by M. L. Fernald (1950). The name Potentilla procumbens Sibthorp was previously used for this species; that is a superfluous and illegitimate name. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
The major part of the range assigned to Potentilla vahliana by E. Hultén (1968) belongs to P. subvahliana. Two morphologic types are present within what is accepted here as Potentilla subvahliana. Plants corresponding to the type of P. subvahliana (Wrangel Island) are widespread throughout northeastern Asia (Chukotka) and arctic North America to northwestern Greenland. Some plants in alpine Alaska, Yukon, and east to Amundsen Gulf, Nunavut, are redder, more densely columnar, and less hairy, with smaller leaves having fewer and narrower lobes, more slender one-flowered stems, narrow and entire bracts, narrower sepals, and much narrower epicalyx bractlets. Potentilla subvahliana is tetraploid, perhaps an allopolyploid, and could consist of lineages from different parental combinations. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 9, p. 135. | FNA vol. 9, p. 203. |
Parent taxa | Rosaceae > subfam. Rosoideae > tribe Potentilleae > Potentilla > sect. Potentilla | Rosaceae > subfam. Rosoideae > tribe Potentilleae > Potentilla > sect. Niveae |
Sibling taxa | ||
Name authority | Laicharding: Veg. Europ. 1: 475. (1790) | Jurtzev: in A. I. Tolmatchew, Fl. Arct. URSS 9(1): 319. (1984) |
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