Potentilla anglica
Potentilla subjuga
English cinquefoil, potentille d'angleterre, trailing cinquefoil, trailing tormentil, wood cinquefoil
Colorado cinquefoil
soon becoming prostrate, ± flagelliform, usually openly branched, eventually rooting at some nodes, (0.3–)1.5–7+ dm.
(0.8–)1–2.5(–3.5) dm.
± persistent, ternate or palmate, 2–10(–12) cm;
petiole 1–7(–8) cm, long hairs sparse to abundant, appressed, 0.5–1 mm, stiff, glands absent;
leaflets 3–5, central ± obovate to cuneate, 1–3(–3.5) × 0.8–2(–2.5) cm, distal ± 1/2 of margin incised 1/4–1/3(–1/2) to midvein, teeth 2–4 per side, surfaces similar, green, sparsely to moderately hairy.
usually palmate with additional lateral leaflets, sometimes pinnate, 3–10(–14) cm;
petiole 1.5–5 cm, vestiture seasonally dimorphic, long hairs abundant, spreading on first-formed leaves, tightly appressed to ascending on later-formed leaves, 1–2 mm, ± stiff (especially on later-formed leaves), cottony and crisped hairs usually absent, glands absent or sparse;
leaflets (3–)5 at tip of leaf axis plus 1(–2) additional pair(s) separated from tip by 3–20 mm, on distal 1/10–1/3(–1/2) of leaf axis, largest leaflets oblanceolate-oblong, (0.5–)1.5–2.5(–3) × 0.3–1 cm, ± whole margin incised 1/2–2/3(–3/4) to midvein, teeth (2–)4–9 per side, usually touching to strongly overlapping, sometimes separate, 2–6 mm, surfaces usually strongly dissimilar (less so on first-formed leaves), abaxial usually white, straight hairs ± abundant (mostly on veins), 1–2 mm, cottony or crisped/cottony hairs ± dense (sparser on first-formed leaves), glands absent or obscured, adaxial green (to grayish), straight hairs sparse to common, 0.5–1.5 mm, cottony and crisped hairs absent, glands sparse.
2–3(–4) proximal to 1st flowering or branching node, usually well expanded at anthesis, usually ternate, 2–6(–10) cm;
petiole 0.3–4(–8) cm;
leaflets (3–)5, ± resembling or narrower than those of basal leaves, narrowly cuneate, 1–2(–3.5) × 0.8–2(–2.5) cm, apex rounded to obtuse.
1–3.
solitary flowers at stolon nodes.
3–20(–30)-flowered.
(1–)3–10(–17) cm.
0.5–2 cm (proximal to 3 cm).
4(–5)-merous;
epicalyx bractlets narrowly elliptic to oblong or ovate, 3–4(–7) × 1–1.5 mm, smaller than to slightly larger than sepals;
hypanthium 2–4 mm diam.;
sepals (3–)4–6 mm, apex broadly acute or acuminate;
petals 6–9 × 5–9 mm, apex ± retuse;
stamens 15–20, filaments 0.8–1.2 mm, anthers 0.8–1.2 mm;
carpels 20–50, styles 0.9–1.5 mm.
epicalyx bractlets narrowly to broadly lanceolate, 2–5(–6) × 1–1.5 mm;
sepals 4–7 mm, apex acute to acuminate;
petals 4–8 × 4–8 mm;
filaments (0.5–)1–2 mm, anthers 0.3–0.8 mm;
carpels 15–30, styles filiform to filiform-tapered, ± papillate-swollen in less than proximal 1/5, 1.5–2 mm.
1–1.5(–1.8) mm, ± smooth.
1.2–1.6 mm.
± erect, stout to slender, 1–4 cm.
= 28, 56 (Europe).
Potentilla anglica
Potentilla subjuga
Potentilla anglica apparently is a product of hybridization involving P. erecta and P. reptans (B. Matfield and J. R. Ellis 1972) that has become stabilized and distinct from both parents. In Europe, it forms back-cross hybrids with both P. erecta (P. ×suberecta Zimmeter) and P. reptans (P. ×mixta Nolte); these hybrids are not known from North America. A garden hybrid supposedly between P. anglica and P. nepalensis Hooker (known as P. ×tonguei Mallett) was found in Allegany State Park, New York, but this was likely cultivated rather than naturalized.
Potentilla anglica probably was introduced even in Newfoundland (A. Kurtto et al. in J. Jalas et al. 1972+, vol. 13), contrary to the view expressed by M. L. Fernald (1950).
The name Potentilla procumbens Sibthorp was previously used for this species; that is a superfluous and illegitimate name.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Potentilla subjuga is centered in the high mountains of Colorado and barely enters New Mexico in the Sangre de Cristo Mountains. Collections from Alberta also apparently belong to this species, but all known collections from Wyoming have been identified as different taxa, at least one currently undescribed. At its most distinctive, P. subjuga is easily recognized by its unique leaf division, with five palmate leaflets subtended by an additional pair (or two) of lateral leaflets. Southern populations, however, are more likely to have only three apical leaflets. The leaflets tend to be strongly bicolored with overlapping teeth, in contrast to most sympatric pinnate species. Petiole vestiture is also distinctive in being seasonally dimorphic, with long hairs on first-formed leaves spreading to ascending and those on later formed leaves tightly appressed, as well as more conspicuously verrucose. Unresolved infraspecific variation exists, and field observations suggest that P. subjuga readily hybridizes with sympatric species, creating a swarm of intermediate specimens.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
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