Potentilla anglica |
Potentilla glaucophylla |
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English cinquefoil, potentille d'angleterre, trailing cinquefoil, trailing tormentil, wood cinquefoil |
blue-leaf cinquefoil, different-leaf cinquefoil, diverse-leaf cinquefoil, mountain meadow cinquefoil, vari-leaf cinquefoil |
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Glands | absent or inconspicuous, uncolored. |
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Stems | soon becoming prostrate, ± flagelliform, usually openly branched, eventually rooting at some nodes, (0.3–)1.5–7+ dm. |
mostly ± ascending, 0.5–3(–4.5) dm. |
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Basal leaves | ± persistent, ternate or palmate, 2–10(–12) cm; petiole 1–7(–8) cm, long hairs sparse to abundant, appressed, 0.5–1 mm, stiff, glands absent; leaflets 3–5, central ± obovate to cuneate, 1–3(–3.5) × 0.8–2(–2.5) cm, distal ± 1/2 of margin incised 1/4–1/3(–1/2) to midvein, teeth 2–4 per side, surfaces similar, green, sparsely to moderately hairy. |
often 2-ranked, palmate, sometimes subpalmate, 2–20 cm; petiole (0.5–)1–8(–14) cm, long hairs often absent, sometimes sparse to abundant, usually appressed, 1–2 mm, weak to ± stiff, short, crisped, and cottony hairs absent, glands absent or sparse; leaflets 5–6(–7), sometimes with 1–2(–4) additional, smaller, entire leaflets, on tip or to distal 1/10(–1/4) of leaf axis, separate, largest ones oblanceolate or cuneate to obovate, 1–4(–6) × 0.5–1.5(–2) cm, margins flat, distal 1/3–1/2(–2/3) ± evenly incised 1/2 or nearly to midvein, undivided medial blade 1.5–7(–10) mm wide, teeth 1–3(–5) per side, broadly lanceolate or linear to narrowly oblong, 1–5(–8) mm, surfaces similar adaxial ± less hairy, ± blue-green, usually glaucous, long hairs sparse to abundant (often restricted to veins, distal teeth, and margins), short, crisped, and cottony hairs absent, glands absent or sparse. |
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Cauline leaves | 2–3(–4) proximal to 1st flowering or branching node, usually well expanded at anthesis, usually ternate, 2–6(–10) cm; petiole 0.3–4(–8) cm; leaflets (3–)5, ± resembling or narrower than those of basal leaves, narrowly cuneate, 1–2(–3.5) × 0.8–2(–2.5) cm, apex rounded to obtuse. |
(0–)1–2(–3). |
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Inflorescences | solitary flowers at stolon nodes. |
2–10(–20)-flowered. |
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Pedicels | (1–)3–10(–17) cm. |
(0.5–)1–3(–6.5) cm. |
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Flowers | 4(–5)-merous; epicalyx bractlets narrowly elliptic to oblong or ovate, 3–4(–7) × 1–1.5 mm, smaller than to slightly larger than sepals; hypanthium 2–4 mm diam.; sepals (3–)4–6 mm, apex broadly acute or acuminate; petals 6–9 × 5–9 mm, apex ± retuse; stamens 15–20, filaments 0.8–1.2 mm, anthers 0.8–1.2 mm; carpels 20–50, styles 0.9–1.5 mm. |
epicalyx bractlets lanceolate to elliptic, 2–5 × 0.8–1.2(–1.6) mm, hairs sparse to common, rarely abundant, ± ascending, glands absent or inconspicuous; hypanthium 3.5–5 mm diam.; sepals (2–)2.5–4.5(–5) mm, apex acute to short acuminate; petals (4–)5–10(–12) × 4–9(–10) mm; filaments 1.5–2.5(–3) mm, anthers 0.4–0.7(–0.9) mm; carpels 25–40, styles filiform above papillate-swollen base, 1.5–2.5(–3) mm. |
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Achenes | 1–1.5(–1.8) mm, ± smooth. |
1.2–1.6 mm. |
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Rootstocks | ± erect, stout to slender, 1–4 cm. |
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2n | = 28, 56 (Europe). |
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Potentilla anglica |
Potentilla glaucophylla |
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Phenology | Flowering May–Aug(–Nov). | |||||
Habitat | Moist flats and slopes, mainly on acidic soil | |||||
Elevation | 0–1200 m (0–3900 ft) | |||||
Distribution |
CA; CO; ME; NY; OR; PA; UT; WA; BC; NL; NS; QC; SPM; Europe [Introduced in North America; introduced also in Atlantic Islands (Azores, Madeira), Pacific Islands (New Zealand), Australia]
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AK; CA; CO; ID; MT; NV; OR; UT; WA; WY; AB; BC; NL; NT; SK; YT
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Discussion | Potentilla anglica apparently is a product of hybridization involving P. erecta and P. reptans (B. Matfield and J. R. Ellis 1972) that has become stabilized and distinct from both parents. In Europe, it forms back-cross hybrids with both P. erecta (P. ×suberecta Zimmeter) and P. reptans (P. ×mixta Nolte); these hybrids are not known from North America. A garden hybrid supposedly between P. anglica and P. nepalensis Hooker (known as P. ×tonguei Mallett) was found in Allegany State Park, New York, but this was likely cultivated rather than naturalized. Potentilla anglica probably was introduced even in Newfoundland (A. Kurtto et al. in J. Jalas et al. 1972+, vol. 13), contrary to the view expressed by M. L. Fernald (1950). The name Potentilla procumbens Sibthorp was previously used for this species; that is a superfluous and illegitimate name. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Varieties 2 (2 in the flora). Potentilla glaucophylla replaces P. diversifolia; an examination of the lectotype of the latter confirms J. Soják’s (1996) conclusion that P. diversifolia applies to a hybrid involving P. glaucophylla and P. hippiana. Historically, this species was often termed P. dissecta Pursh, a misapplied name that has been rejected (B. Ertter et al. 2008). In addition to the nomenclatural change, the circumscription of Potentilla glaucophylla is here restricted to plants with usually glaucous, blue-green, distally toothed leaflets that are usually glabrate (at least in var. glaucophylla). Plants from the Colorado Plateau and southern Rocky Mountains formerly included in this species and having dark green leaves, more leaflet teeth, and larger anthers are transferred either to P. townsendii or to a currently undescribed entity. The two-ranked leaves of Potentilla glaucophylla suggest a possible connection to P. crantzii in sect. Aureae. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 9, p. 135. | FNA vol. 9, p. 152. | ||||
Parent taxa | Rosaceae > subfam. Rosoideae > tribe Potentilleae > Potentilla > sect. Potentilla | Rosaceae > subfam. Rosoideae > tribe Potentilleae > Potentilla > sect. Graciles | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | P. diversifolia var. glaucophylla | |||||
Name authority | Laicharding: Veg. Europ. 1: 475. (1790) | Lehmann: Index Seminum (Hamburg) 1836: 7. (1836) | ||||
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