Potentilla anglica
Potentilla brunnescens
English cinquefoil, potentille d'angleterre, trailing cinquefoil, trailing tormentil, wood cinquefoil
brownish cinquefoil, slender brown cinquefoil, slender cinquefoil
conspicuous, usually golden.
soon becoming prostrate, ± flagelliform, usually openly branched, eventually rooting at some nodes, (0.3–)1.5–7+ dm.
ascending to nearly erect, (1.5–)3–7 dm.
± persistent, ternate or palmate, 2–10(–12) cm;
petiole 1–7(–8) cm, long hairs sparse to abundant, appressed, 0.5–1 mm, stiff, glands absent;
leaflets 3–5, central ± obovate to cuneate, 1–3(–3.5) × 0.8–2(–2.5) cm, distal ± 1/2 of margin incised 1/4–1/3(–1/2) to midvein, teeth 2–4 per side, surfaces similar, green, sparsely to moderately hairy.
sometimes 2-ranked, palmate, (5–)10–30 cm;
petiole (1.5–)5–25(–50) cm, long hairs sparse to abundant, appressed to spreading, 1–2 mm, ± stiff, short, crisped, and cottony hairs absent, glands ± abundant;
leaflets (5–)7–9, at tip of leaf axis, separate to slightly overlapping, largest ones oblanceolate-elliptic to narrowly obovate, (2–)5–10 × 1–4 cm, margins flat to ± revolute, distal 3/4 to nearly whole length ± evenly incised (1/4–)1/2–3/4+ to midvein, undivided medial blade 2–15(–20) mm wide, teeth (5–)7–10 per side, linear to broadly lanceolate, 3–15 mm, surfaces ± similar, abaxial paler, green, not glaucous, long hairs sparse to common, short-crisped hairs absent or sparse, cottony hairs absent, glands ± abundant.
2–3(–4) proximal to 1st flowering or branching node, usually well expanded at anthesis, usually ternate, 2–6(–10) cm;
petiole 0.3–4(–8) cm;
leaflets (3–)5, ± resembling or narrower than those of basal leaves, narrowly cuneate, 1–2(–3.5) × 0.8–2(–2.5) cm, apex rounded to obtuse.
1–3(–4).
solitary flowers at stolon nodes.
12–30(–70)-flowered.
(1–)3–10(–17) cm.
0.2–1(–4) cm.
4(–5)-merous;
epicalyx bractlets narrowly elliptic to oblong or ovate, 3–4(–7) × 1–1.5 mm, smaller than to slightly larger than sepals;
hypanthium 2–4 mm diam.;
sepals (3–)4–6 mm, apex broadly acute or acuminate;
petals 6–9 × 5–9 mm, apex ± retuse;
stamens 15–20, filaments 0.8–1.2 mm, anthers 0.8–1.2 mm;
carpels 20–50, styles 0.9–1.5 mm.
epicalyx bractlets lanceolate to elliptic, 3–6 × 1–2 mm, hairs sparse to common, loosely appressed to spreading, glands common to abundant;
hypanthium 4–6(–8) mm diam.;
sepals 4–8(–9) mm, apex acute to acuminate;
petals 6–9 × 6–10 mm;
filaments 1–2 mm, anthers 0.6–1 mm;
carpels 30–60, styles filiform-tapered, papillate-swollen proximally, 1.5–2 mm.
1–1.5(–1.8) mm, ± smooth.
1.5 mm.
± erect, stout to slender, 1–4 cm.
= 28, 56 (Europe).
Potentilla anglica
Potentilla brunnescens
Potentilla anglica apparently is a product of hybridization involving P. erecta and P. reptans (B. Matfield and J. R. Ellis 1972) that has become stabilized and distinct from both parents. In Europe, it forms back-cross hybrids with both P. erecta (P. ×suberecta Zimmeter) and P. reptans (P. ×mixta Nolte); these hybrids are not known from North America. A garden hybrid supposedly between P. anglica and P. nepalensis Hooker (known as P. ×tonguei Mallett) was found in Allegany State Park, New York, but this was likely cultivated rather than naturalized.
Potentilla anglica probably was introduced even in Newfoundland (A. Kurtto et al. in J. Jalas et al. 1972+, vol. 13), contrary to the view expressed by M. L. Fernald (1950).
The name Potentilla procumbens Sibthorp was previously used for this species; that is a superfluous and illegitimate name.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
The conspicuous golden glands, dark green color, and relatively broad leaflets make Potentilla brunnescens one of the better-defined species of sect. Graciles, though it has often been treated as a variety of P. gracilis. The glands, which are often on septate stalks, are particularly conspicuous on the epicalyx bractlets, hypanthium, and sepals, and at the junction of petiole and leaflets. The species occurs from the Blue Mountains of Oregon and Washington to western Montana, northern Utah, and north-central Colorado; collections from Alaska are probably not native.
As presented here, Potentilla brunnescens encompasses two morphological extremes that might merit varietal status. Leaflets of the more common and widespread extreme are incised about halfway to the midvein; in contrast, some populations in western Wyoming and adjacent Idaho have leaflets incised 3/4 or more to the midvein. The latter have often been assigned to P. gracilis var. elmeri while reserving P. gracilis var. brunnescens for the former; this is untenable because the type of P. brunnescens is the latter form.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
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