Polytrichum strictum |
Polytrichum commune |
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bog haircap moss, hummock haircap, narrow-leaf haircap, polytrichum moss, slender haircap |
common haircap moss, polytrichum moss |
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Habit | Plants slender, green to whitish green, dark brownish with age, in deep, compact tufts. | Plants medium to robust, dark green to brownish with age, in loose or rather dense, tall tufts, often in extensive patches. | ||||
Stems | 6–12(–20) cm, simple, densely matted with wooly whitish to light-brownish tomentum. |
(2–)5–10(–70) cm, stiffly erect to decumbent, mostly simple or rarely forked, bracteate proximally, densely to rather distantly leafy. |
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Leaves | 2–5(–6) mm, erect to closely appressed when dry, erect-spreading when moist; sheath oblong-rectangular, brownish, ± abruptly contracted to the blade; blade narrowly lanceolate, acuminate, flat, with sharply infolded margins; marginal lamina 6–7 cells wide, 1-stratose, entire to finely crenulate above, membranous and transparent, abruptly infolded and enclosing the lamellae and overlapping towards the apex; costa toothed abaxially towards the apex, short-excurrent as a short, reddish brown awn; lamellae bluntly crenate in profile, 5–8 cells high, the marginal cells in section pyriform, thick-walled, ending in a thickened knob, end cells of lateral lamellae ovoid and scarcely thickened at the apex; sheath cells 45–80 × 7–10 µm, elongate-rectangular (5–7:1), narrower toward the margin; cells of the marginal lamina transversely elongate, shorter and obliquely oriented towards the margins, very thick-walled and colorless. |
6–8(–12) mm, erect or erect-spreading when dry, flexuose, with distinctly recurved tips, spreading to broadly recurved when moist, or the blade sharply squarrose-reflexed from the base; sheath oblong to elliptic, involute-tubular and clasping the stem, often golden yellow and highly nitid, abruptly contracted to the narrowly lanceolate blade; marginal lamina plane or erect, narrow, 2–3(–7) cells wide, toothed from the base of blade to the apex, the teeth unicellular, embedded in the margin; costa toothed abaxially near the apex, excurrent, ending in short, rough awn; lamellae crenulate in profile, 5–9 cells high, the margin distinctly grooved as seen from above, with 2 rows of paired, projecting knobs, the marginal cells in section narrow or more typically enlarged and wider than those beneath, retuse to deeply notched (rarely divided by a vertical partition), ± thick-walled and brownish, smooth; sheath cells 60–90 × 10–13 µm, elongate-rectangular to linear (as much as 20:1), narrower toward the margins; cells of the marginal lamina 10–15 µm wide, subquadrate. |
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Seta | 2–4 cm, yellowish to reddish brown. |
5–9 cm, stout, yellowish to reddish brown. |
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Sexual condition | dioicous; perichaetial leaves somewhat longer than the stem leaves, ending in a slender awn. |
dioicous; perichaetial leaves with a long, scarious-margined sheath and blade greatly reduced, gradually narrowed to a finely acuminate tip, the margins toothed, denticulate to subentire, the costa short- to long-excurrent, roughened to almost smooth. |
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Capsule | 2–3 mm, short-rectangular to almost cubic (1–1.5:1), brownish, sharply 4-angled and prismatic, suberect, becoming horizontal when ripe; peristome 200–230 µm, divided to 0.8, the teeth 64, obtuse. |
3–6 mm, short-rectangular to cubic, brown to dark reddish brown, glaucous when fresh, sharply 4-alate, inclined to horizontal; peristome 250 µm, divided to 0.6, the teeth 64, pale. |
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Calyptra | dirty white to light brown, enclosing the capsule. |
golden yellow to brownish, completely enveloping the capsule. |
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Spores | 7–9(–15) µm. |
5–8(–12) µm. |
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Polytrichum strictum |
Polytrichum commune |
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Habitat | Sphagnum bogs, wet heaths and tundra, muskeg, sedge meadows, moist alpine tundra, also on local elevations and on rotten stumps in wet spruce forests | |||||
Elevation | low to high elevations | |||||
Distribution |
AK; CA; CO; CT; GA; IA; IL; IN; MA; MD; ME; MI; MN; MT; NC; NH; NJ; NY; OH; OR; PA; RI; UT; VT; WA; WI; WV; WY; AB; BC; MB; NB; NL; NS; NT; NU; ON; PE; QC; SK; YT; South America; e Asia (Russia, Japan); Atlantic Islands (Faroes, Iceland); Greenland; n Asia; n Europe (Scandinavia, Svalbard); Antarctica
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Mexico; Widely distributed in the North America; throughout temperate and boreal latitudes in the Northern Hemisphere; Pacific Islands (New Zealand); Australia
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Discussion | Polytrichum strictum is widespread in the boreal regions of the Holarctic, and is one of the commonest low arctic representatives of the family (D. G. Long 1985), with survivals southward in relict bogs, for example in northern Indiana, northern Illinois, and northwestern Iowa, also in alpine situations in the eastern mountains to the Carolinas and Georgia. In Nunavut, it is known from Baffin, Bathurst, and Devon islands. Its characteristic habitat is on hummocks in Sphagnum bogs, in deep masses tightly bound together by dirty-white, wooly tomentum, with short, stiffly erect leaves, and cubical capsules, a clear correlation between a distinctive morphology, distribution, and ecology. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Widely distributed in the North America, throughout temperate and boreal latitudes in the Northern Hemisphere, Mexico, Pacific Islands (New Zealand), Australia. The only North American species likely to be confused with Polytrichum commune is Polytrichastrum pallidisetum. When sterile, Polytrichum commune is easily distinguished by cross-sections of the leaves, particularly by the greatly broadened costa, and the notched marginal cells of the lamellae. In Polytrichastrum pallidisetum the marginal cells of the lamellae are broadened and shallowly retuse, but not distinctly notched, and in Polytrichum commune the arc of guide cells (seen in section below mid-point of blade) is broad, 18–24 cells wide, compared with 8–12 cells wide in Polytrichastrum pallidisetum. When capsules are present, the two species can be distinguished at a glance. A striking form of Polytrichum commune growing in wetlands, var. uliginosum, has greatly elongate stems and tightly sheathing, glossy leaf bases, the blades are flexuose and squarrose-recurved. Recent European genetic studies (R. J. Bijlsma et al. 2000) suggest that it should be recognized as a separate species. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 27, p. 139. | FNA vol. 27, p. 135. | ||||
Parent taxa | Polytrichaceae > Polytrichum | Polytrichaceae > Polytrichum | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | P. affine, P. juniperinum var. affine, P. juniperinum var. gracilius | |||||
Name authority | Bridel: J. Bot. (Schrader) 1800(1): 286. (1801) | Hedwig: Sp. Musc. Frond., 88. (1801) | ||||
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