Polytrichum piliferum |
Polytrichum swartzii |
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awn haircap moss, bristly haircap moss, polytrichum moss |
swartz's polytrichum moss |
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Habit | Plants small to medium, glaucous green to reddish brown, in loose tufts. | Plants often rather soft and flexuose, green to blackish when old. |
Stems | (0.5–)1–4 cm tall, rather wiry, unbranched, comose at the tips, whitish tomentose only near the base. |
2–9 cm, simple, erect, in proximal part moderately to densely brownish tomentose. |
Leaves | (2–)3–4 mm, erect, straight and slightly incurved when dry, erect-spreading when moist; sheath ovate, ± contracted to the blade; blade linear-lanceolate, turgid, with sharply infolded margins, the leaf apex abruptly contracted to the base of the awn; marginal lamina 5–8 cells wide, 1-stratose, membranous, entire to finely serrulate toward the apex, enclosing the lamellae and overlapping in distal half or more; costa typically smooth abaxially, long-excurrent as a spinulose-toothed, hyaline awn; lamellae in profile crenulate-dentate to serrulate, with crenulations directed towards the leaf apex, (4–)6–8 cells high, the marginal cells in section conic to distinctly pyriform, terminating in a distinct knob, the marginal cells of the lateral lamellae ovoid, thinner-walled; sheath cells 60–80 × 10–15 µm, elongate-rectangular (4–6:1); cells of marginal lamina transversely elongated, ± irregular and sinuous, smaller toward the margins and obliquely oriented, especially approaching the apex, thick-walled. |
3–8 mm, loosely imbricate, appressed to erect-spreading and flexuose when dry, patent to widely spreading and weakly recurved when moist; sheath rectangular, scarcely narrowed to the blade; blade lanceolate to linear-lanceolate, often caducous at the junction of sheath and blade, the apex subulate, weakly channeled; marginal lamina 6–9 cells wide, distantly toothed to subentire; costa excurrent as short brown entire to serrulate tip, smooth abaxially or with a few teeth near apex; lamellae in profile entire to shallowly crenulate, 5–10 cells high, the marginal cells in section usually somewhat broadened, flat-topped or shallowly grooved, single or geminate, thin-walled, smooth, the marginal cells of lateral lamellae asymmetric; median sheath cells 75–110 × 2–12 µm, linear; cells of marginal lamina 9–15 µm, quadrate, thin- to firm-walled; perichaetial leaves with long sheathing bases and short subulate blade. |
Seta | 1–3 cm, stout, flexuose, reddish brown. |
2.5–5 cm, reddish brown. |
Sexual condition | dioicous; perigonia intense wine-red; perichaetial leaves a little longer than foliage leaves, with hyaline lamina and longer awns. |
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Capsule | 2.5–3.5 mm, short to almost cubic (1.5–2:1), inclined, becoming horizontal when mature; peristome rather short, 110–180(–200) µm, divided to 0.6, the teeth about 64, obtuse. |
2.5–3 cm, ± cubic, sharply 4–angled, suberect when mature, becoming horizontal when old; peristome teeth 64, 160–210 µm, obtuse, the basal portion 60–75 µm. |
Calyptra | dirty white to light brown, enclosing the capsule. |
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Spores | 9–12 µm. |
12–15 µm. |
Polytrichum piliferum |
Polytrichum swartzii |
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Habitat | Shallow well-drained sandy or gravelly soil over rocks and boulders in sunny situations, often associated with Cladonia, in road cuts, old fields, burned over areas, heaths, rocky ridges and moraines, and in dry alpine tundra and late snow areas northward | Very wet and regularly flooded situations, sedge meadows, wet tundra and lake shores (D. G. Long 1985) |
Elevation | low to moderate elevations | |
Distribution |
AK; AL; AZ; CA; CO; CT; IA; ID; IN; KS; KY; LA; MA; ME; MI; MN; MO; MT; NC; ND; NE; NH; NJ; NM; NY; OH; OR; PA; SD; TN; UT; VA; VT; WA; WI; WV; WY; HI; AB; BC; MB; NB; NL; NS; NT; NU; ON; PE; QC; YT; South America; Australia; Greenland; Atlantic Islands (Falkland Islands, Macaronesia); Asia; Europe; Antarctica
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AK; LB; NT; NU; YT; Greenland; Europe (Scandinavia); n Asia; e Asia; Atlantic Islands (Iceland) |
Discussion | Polytrichum piliferum is easily recognized by its habit, growing in short, loose reddish brown clumps, each plant crowned by a whitish tuft of intertwined hyaline awns. The intensely colored, wine-red antheridial rosettes are remarkably flower-like in appearance. The lamella marginal cells are pyriform in section, ending in a distinct knob, except for the lamellae standing in the shade of the inflexed lamina. The leaves of P. piliferum are smooth abaxially and abruptly contracted to the base of the awn; P. juniperinum is a larger plant with reddish rather than hyaline awns; P. hyperboreum is typically fastigiately branched, the leaves with a channeled leaf apex, tapering to a bicolored awn. In Nunavut, it is known from Baffin and Melville islands. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Polytrichum swartzii is a northern species with distantly toothed to subentire leaves, differing chiefly by the rounded-quadrate, flat-topped (not retuse or grooved) and scarcely-thickened marginal cells of the lamellae. The capsules are shortly cubic. In Nunavut, it is known from Baffin and Devon islands. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 27, p. 140. | FNA vol. 27, p. 137. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | P. commune var. swartzii, P. algidum, P. inconstans | |
Name authority | Hedwig: Sp. Musc. Frond., 90. (1801) | C. J. Hartman: Handb. Skand. Fl. ed. 5, 361. (1849) |
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