Polystichum imbricans |
Polystichum scopulinum |
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imbricate sword-fern, narrow-leaf sword fern, rock sword-fern |
Eaton's holly fern, mountain holly fern, polystic des rochers, rock sword fern |
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Stems | ascending to erect. |
ascending. |
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Leaves | erect to arching back at tip, 2–8 dm; bulblets absent. |
erect, 1–3(–5) dm; bulblets absent. |
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Petiole | 1/4–1/3 length of leaf; scales abruptly diminishing in size distally and falling off early but retaining conspicuous tuft of brown scales at base. |
1/5–1/3 length of leaf, densely scaly but scales falling off distally; scales light brown, abruptly diminishing in size distally. |
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Blade | linear-lanceolate to linear, 1-pinnate, base not or slightly narrowed. |
narrowly lanceolate, 1-pinnate-pinnatifid, base narrowed. |
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Pinnae | oblong, slenderly lanceolate, or falcate, usually overlapping, in 1 plane or twisted out of plane of blade, 2–4 cm; base oblique, auricles well developed; margins serrulate-spiny with teeth ascending; apex cuspidate or apiculate with subapical teeth smaller than apical tooth; microscales lanceolate to linear with straight or sharply angular projections, sparse, on abaxial surface only. |
oblong-lanceolate, overlapping, folded inward and twisted horizontally, 1–3 cm; base oblique; margins serrulate with teeth curved inward; apex obtuse to cuspidate with subapical teeth smaller than apical tooth; microscales narrowly lanceolate, with stout projections, sparse, on abaxial surface only. |
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Indusia | entire to sharply dentate. |
entire-ciliate. |
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Spores | dark brown. |
brown. |
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2n | = 164. |
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Polystichum imbricans |
Polystichum scopulinum |
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Habitat | Rock crevices and at base of boulders, serpentine to acidic substrates, usually exposed to full sun | |||||
Elevation | 0–3500 m (0–11500 ft) | |||||
Distribution |
CA; OR; WA; BC; only in the flora
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AZ; CA; CO; ID; MT; NV; OR; UT; WA; WY; BC; NF; QC
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Discussion | Polystichum imbricans is one of the postulated ancestors of two allopolyploids, P. californicum and P. scopulinum (D. H. Wagner 1979). Relationships to P. munitum are discussed under that species. Subspecies 2. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Polystichum scopulinum is widely distributed in the United States west of the 110th meridian, where it occurs in sporadic, usually small populations. The species is abundant only on montane serpentine outcrops. The populations in Newfoundland and Quebec are dramatically disjunct. Polystichum scopulinum is an allopolyploid, believed on morphologic grounds to be derived from P. imbricans × lemmonii (D. H. Wagner 1979). Based on putative hybridization between P. scopulinum and P. munitum (P. S. Soltis et al. 1989; W. H. Wagner Jr. 1973), however, P. munitum may also be involved. This hybrid is discussed under P. californicum. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 2. | FNA vol. 2. | ||||
Parent taxa | Dryopteridaceae > Polystichum | Dryopteridaceae > Polystichum | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | Aspidium munitum var. imbricans, P. munitum subsp. imbricans, P. munitum var. imbricans | Aspidium aculeatum var. scopulinum, P. mohrioides var. scopulinum | ||||
Name authority | (D. C. Eaton) D. H. Wagner: Pteridologia 1: 50. (1979) | (D. C. Eaton) Maxon: Fern Bull. 8: 29. (1900) | ||||
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