Polystichum imbricans |
Polystichum lemmonii |
|||||
---|---|---|---|---|---|---|
imbricate sword-fern, narrow-leaf sword fern, rock sword-fern |
Lemmon's holly fern, Lemmon's sword fern, Shasta fern, Shasta sword fern |
|||||
Stems | ascending to erect. |
decumbent to ascending. |
||||
Leaves | erect to arching back at tip, 2–8 dm; bulblets absent. |
erect, 1–3.5 dm; bulblets absent. |
||||
Petiole | 1/4–1/3 length of leaf; scales abruptly diminishing in size distally and falling off early but retaining conspicuous tuft of brown scales at base. |
1/5–1/4 length of leaf, sparsely scaly; scales pale tan, abruptly diminishing in size distally. |
||||
Blade | linear-lanceolate to linear, 1-pinnate, base not or slightly narrowed. |
narrowly lanceolate, 2-pinnate, scarcely narrowed at base. |
||||
Pinnae | oblong, slenderly lanceolate, or falcate, usually overlapping, in 1 plane or twisted out of plane of blade, 2–4 cm; base oblique, auricles well developed; margins serrulate-spiny with teeth ascending; apex cuspidate or apiculate with subapical teeth smaller than apical tooth; microscales lanceolate to linear with straight or sharply angular projections, sparse, on abaxial surface only. |
ovate, overlapping, folded inward and twisted horizontally, 0.5–2 cm; base truncate to oblique, proximal acroscopic pinnules not enlarged; apex broadly acute; microscales narrowly lanceolate, with few projections, sparse, ± confined to costa of both surfaces. |
||||
Indusia | entire to sharply dentate. |
entire or minutely dentate-erose. |
||||
Spores | dark brown. |
dark brown to blackish. |
||||
Pinnules | ± stalked, rounded, acroscopic auricle not well developed, margins entire to weakly dentate, apex rounded. |
|||||
2n | = 82. |
|||||
Polystichum imbricans |
Polystichum lemmonii |
|||||
Habitat | On rocky serpentine slopes | |||||
Elevation | 1200–2400 m (3900–7900 ft) | |||||
Distribution |
CA; OR; WA; BC; only in the flora
|
CA; OR; WA; BC
|
||||
Discussion | Polystichum imbricans is one of the postulated ancestors of two allopolyploids, P. californicum and P. scopulinum (D. H. Wagner 1979). Relationships to P. munitum are discussed under that species. Subspecies 2. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Polystichum lemmonii forms sterile hybrids with P. scopulinum and P. munitum. The first hybrid may be abundant where the two parents grow together, which they frequently do in the Wenatchee Mountains of Washington and Siskiyou Mountains of northern California and southwest Oregon. The hybrid is very similar to P. lemmonii but has malformed sporangia and slightly less divided pinnae than P. lemmonii. The P. lemmonii × P. munitum hybrid is morphologically indistinguishable from P. scopulinum; it is a sterile diploid reported only twice from the Wenatchee Mountains of Washington (W. H. Wagner Jr. 1973; P. S. Soltis et al. 1989). It is possible that this hybrid involves P. imbricans and not P. munitum; neither study distinguished between them. American authors have misapplied the name Polystichum mohrioides (Bory) C. Presl, a South American species, to P. lemmonii. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
||||
Key |
|
|||||
Source | FNA vol. 2. | FNA vol. 2. | ||||
Parent taxa | Dryopteridaceae > Polystichum | Dryopteridaceae > Polystichum | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | Aspidium munitum var. imbricans, P. munitum subsp. imbricans, P. munitum var. imbricans | P. mohrioides var. lemmonii | ||||
Name authority | (D. C. Eaton) D. H. Wagner: Pteridologia 1: 50. (1979) | L. Underwood: Native Ferns ed. 6 116. (1900) | ||||
Web links |
|
|